Studies of how gender is characterized, performed, and understood in outdoor activities in relation to skill development are limited, but growing. This research explored gender and social relationships across levels of recreation specialization in fly-fishing among anglers in Prince George, BC. Semi-structured interviews were conducted with 23 participants, 11 of whom then attended a level-specific fishing day during which participant observation was conducted. Interviews were transcribed and coded. Four main findings were derived. First, anglers’ social relationships shifted from dependence on others to intentional self-expression. Second, anglers learned to belong ecologically and socially through skill development and equipment use. Third, anglers’ relationships with fish moved from possession to communion. Fourth, anglers’ described escaping their daily routines to engage with rural places and fishing. The analysis and discussion show how masculinity was constructed and performed, and highlight the roles of socialization, behaviors, and equipment in shaping and gendering rural settings.