ee eee seven shoveller, four of the last probably being transients, were observed at different times in the region adjacent to Quesnel and west of that place None of these species was recorded in more northern latitudes. Redhead and ruddy duck were absent and only five canvas back. 4 males and 1 female, were observed during the entire season. Coots were conspicuously absent on most marshes and represented by only a few pairs on others. The absence of redhead, canvas-back, ruddy duck, the absence of blue-winged teal and cinnamon teal except in the most southern parts, and the rarity of the coot might be explained by the fact that the region generally is beyond the range-periphery of the Species as developed west of the Rocky Mountains. What limits their expansion in space cannot be explained by the presence or absence of any known physical factors. Perhaps it may be attri- buted to physical elements or limitations inherent in the species themselves. In contrast it can be observed how the mallard has expanded over the continent from California to Alaska. The absence of pintail and shoveller (species that nest in Alaska and the Yukon) as nesting elements in the population is even less readily accounted for. It might be suggested that duck populations within more or less definite geographical boundaries are static in that they do not expand beyond those boundaries. -The size of the population rises and falls in response to condi- tions of water and food, and in response to other factors that may be favourable in some years and unfavourable in others. Natives of any given area and their increase return to the same area each year (the recovery of banded mallards and pintails shows that units of population travel together in successive years), and if conditions are unfavourable some may not breed, or nest unsuccess-— fully, but nevertheless few or none seek more favourable condi- tions outside the boundaries of the region to which they are native. Certain lakes and marshes of central British Columbia which are relatively barren of waterfowl in summer attract large numbers of ducks and geese during migrations. Many observers have reported this in respect of particular waters such as Bulkley Lake, Francois Lake and the Nechako River. Nesting grounds north of the region explored, in which river margins figure prominently, are reported by travellers as populated chiefly by Canada goose, mallard, and American merganser. Finally in summary it may be said that one of the most interest— ing facts brought out by this investigation is that many waters in central British Columbia suited to waterfowl propagation have in- significant populations or are entirely devoid of waterfowl. - 45 -