BIOPHYSICAL DIVERSITY ON LANDSLIDES IN THE PEACE RIVER REGION OF BRITISH COLUMBIA by Victoria Kress B.Sc. NRM, Forestry and Wildlife, University of Northern British Columbia, 2000 MF, University of British Columbia, 2014 THESIS SUBMITTED IN PARTIAL FULFILLMENT OF THE REQUIREMENTS FOR THE DEGREE OF DOCTOR OF PHILOSOPHY IN NATURAL RESOURCES AND ENVIRONMENTAL STUDIES UNIVERSITY OF NORTHERN BRITISH COLUMBIA December 2023 ©Victoria Kress, 2023 ABSTRACT The aim of this dissertation is to investigate and quantify biophysical diversity on landslides in the boreal forest of the Peace River Region of northern British Columbia (BC), sampling three landslides that occurred in the last 50 years. Landslides are increasing in the boreal forest, likely driven by climate changes such as increased precipitation and permafrost thaw, and as a derivative of large wildfires. An understanding of ecosystem recovery on landslides is important for conservation and management purposes. Several studies have been done in southern parts of the world to elucidate processes of landslide recovery. However, few studies have addressed landslide recovery in northern climates, and little is known about the biophysical diversity of landslides in this region. This research investigates whether landslides are more biophysically diverse than the surrounding relatively undisturbed terrain, and whether microsite variables or geomorphic diversity are predictors of plant community diversity. Using a series of field sampling campaigns and GIS (geographic information system) mapping exercises, I show that landslides are more biophysically diverse than the surrounding terrain in some respects, while the surrounding undisturbed terrain is more diverse in other aspects. The age and size of landslides also appear to influence diversity. Microsite diversity does not necessarily predict plant diversity. The research highlights the role of invasive plant species in slope stabilisation and plant community makeup. I also show that landslide ponds are disproportionately concentrated on rotational landslides, and that most ponds on landslides occur in the body and toe. I note post-slide modifications such as drainage of landslide ponds and lowering of landslide ridges, but many geomorphic features are expected to endure for decades to millennia. Overall, the research shows that vegetation recovery is complex and may take decades to fully manifest. This study contributes knowledge about plant community and site diversity on landslides by providing quantitative data and comparing those traits with those found ii on surrounding terrain. These findings can be used as guidance when identifying conservation and management practices for ecological restoration of disturbed slopes. iii TABLE OF CONTENTS ABSTRACT ......................................................................................................................... ii TABLE OF CONTENTS ..................................................................................................... iv LIST OF TABLES ............................................................................................................... vi LIST OF FIGURES ........................................................................................................... viii ACKNOWLEDGEMENTS ............................................................................................... xiii DEDICATION................................................................................................................... xiv Chapter 1. Introduction to landslide biophysical diversity research ....................................... 1 1.1 Introduction................................................................................................................. 1 1.2 Background to the study .............................................................................................. 2 1.3 The research problem .................................................................................................. 6 1.4 Research aims, objectives, and questions ..................................................................... 8 1.5 Significance .............................................................................................................. 13 1.6 Structural outline of the dissertation .......................................................................... 13 1.7 References ................................................................................................................ 15 Chapter 2. Biophysical alpha diversity of landslides and comparison with surrounding undisturbed terrain.............................................................................................................. 18 2.1 Introduction and Background .................................................................................... 18 2.2 Study areas ................................................................................................................ 27 2.3 Methods and Analysis ............................................................................................... 31 2.4 Results ...................................................................................................................... 46 2.5 Discussion ................................................................................................................. 81 2.6 Conclusions and recommendations .......................................................................... 106 2.7 References .............................................................................................................. 113 Chapter 3. Beta diversity as a measure of biophysical turnover on landslides .................... 122 3.1 Introduction and Background .................................................................................. 122 3.2 Methods .................................................................................................................. 127 3.3 Results .................................................................................................................... 131 3.4 Discussion ............................................................................................................... 141 3.5 Conclusions and Recommendations ........................................................................ 159 3.6 References .............................................................................................................. 162 iv Chapter 4. Landslide-generated ponds in the Fort St. John area, British Columbia: Characteristics, distribution, and ecological implications .................................................. 165 4.1. Introduction............................................................................................................ 165 4.2. Study area .............................................................................................................. 168 4.3. Methods ................................................................................................................. 170 4.4. Results ................................................................................................................... 177 4.5. Discussion .............................................................................................................. 193 4.5. Conclusions and recommendations ......................................................................... 203 4.6 References .............................................................................................................. 205 Chapter 5. Conclusions and Recommendations for Landslide Recovery and Management 208 5.1 Conclusions............................................................................................................. 208 5.2 Recommendations ................................................................................................... 212 5.3 Final thoughts ......................................................................................................... 215 APPENDICES.................................................................................................................. 217 Appendix 1 Material Origin classes .................................................................................. 218 Appendix 2 Vegetation summary tables - Mean species cover (% of total relevé area) ...... 219 Appendix 3 Vegetation cover by growth form ................................................................... 236 Appendix 4 Landslide geotypes glossary .......................................................................... 256 Appendix 5 Geotype polygons .......................................................................................... 259 Appendix 6 Microtopography elevation summary tables - BEC plots................................ 274 Appendix 7 Landslide pond descriptions - Mapsheet 94A ................................................. 278 Appendix 8 Pond size statistics by landslide type.............................................................. 302 Appendix 9 Pond size by geomorphic location per slide type ............................................ 303 v LIST OF TABLES Table 2-1. Ten most abundant plant species for relevés at all study sites, comparing landslide and undisturbed results. Values are mean percent cover of the entire relevé area +/- standard error of the mean. 48 Table 2-2. Relevé mean growth form cover as a percentage of mean total vegetation cover, along with proportions, for all study sites. The table provides a comparison between landslide and undisturbed vegetation growth form cover. 51 Table 2-3. Combined* and mean (+/- standard error) vegetation diversity indices for relevés 52 Table 2-4. Hill numbers for vegetation diversity on relevés. 53 Table 2-5. Pearson correlations of BEC plot plant species abundance with NMS Axis 1. N = 116. 61 Table 2-6. Pearson correlations of BEC plot plant species abundance with NMS Axis 2. N = 116. 61 Table 2-7. Pearson correlations of environment variables and Axis 1 and Axis 2 of the ordination space. N=116. 62 Table 2-8. Site types/series found on all landslides and undisturbed terrain in the study. 63 Table 2-9. Diversity indices for site series polygons - all study sites. BE = Beatton River, CE = Cecil Lake, HA = Hasler Flats. 69 Table 2-10. Landslide geotype summary by area (m2) and percentage. 73 Table 2-11. Geotype diversity indices for landslide geotype polygons. 74 Table 2-12. Undisturbed geotype summary by area (m2) and percentage. 75 Table 2-13. Geotype diversity indices for undisturbed geotype polygons. 76 Table 2-14. BEC 50m2 plot LiDAR points: Elevation coefficient of variation (CV) summary statistics. 78 Table 3-1. Landslide transect vegetation beta diversity (Bw) values. 132 Table 3-2. Landslide transect vegetation alpha diversity statistics. 133 Table 3-3. Undisturbed transect vegetation beta diversity (Bw) values. 134 vi Table 3-4. Undisturbed transect vegetation alpha diversity statistics. 135 Table 3-5. Landslide transect environment beta diversity (Bw) values. 136 Table 3-6. Landslide transect environment alpha diversity statistics. 137 Table 3-7. Undisturbed transect environment beta diversity (Bw) values. 138 Table 3-8. Undisturbed transect environment alpha diversity statistics. 139 Table 4-1. Summary information on ponds and landslides in the 768.25 km2 (76,825 ha) study area. 179 vii LIST OF FIGURES Figure 1-1. Beatton River landslide. Top image is an oblique 3D presentation of an ortho mosaic draped over drone imagery (flown 2016), at vertical exaggeration 1X. Bottom image is an aerial view of the landslide, illustrating temporary partial blockage of the Beatton River. (Aerial photo on bottom was taken October 20, 2015, by M. Geertsema. Used with permission.) 10 Figure 1-2. Cecil Lake landslide. Top image is an oblique 3D presentation of World Imagery layer drape over LiDAR imagery, vertical exaggeration 1X. Inset in top image is a photograph of the headscarp and part of the landslide, facing west. Inset photo was taken May 12, 2016. Bottom image is an ortho mosaic presentation of the landslide, showing variable topography and ponds. 11 Figure 1-3. Hasler Flats landslide. Top image is an ortho mosaic draped over drone imagery, vertical exaggeration 1X. Bottom image is an aerial view of the landslide, facing south. (Aerial photo on bottom was taken September 30, 2013, by M. Geertsema. Used with permission.) 12 Figure 2-1. Study areas of Beatton River, Cecil Lake, and Hasler Flats landslides, shown in Google Earth Pro. The landslides are in the Peace River Region of northeastern British Columbia, Canada. Landslide locations are outlined in yellow and identified with red locator balloons. Inset map shows landslide locations at a provincial scale and in relation to Mapsheet 94A, which is outlined in red. BE = Beatton River landslide, CE = Cecil Lake landslide, and HA = Hasler Flats landslide. 29 Figure 2-2. Vegetation and site diversity on landslides in the study. From top left clockwise: horst and graben complex and mature rafts (Hasler Flats. Photo July 12, 2018); unvegetated weathered pillar next to heavy brush (Cecil Lake. Photo September 17, 2017); dewatered/revegetated pond site (Cecil Lake. Photo August 23, 2018); steep unvegetated scarps interspersed with relatively level swaths of abundant invasive vegetation and the occasional pond (Beatton River. Photo August 15, 2017). 47 Figure 2-3. Comparison of rank abundance distributions for relevé plot plant species for all study sites. Abundance rank by species is on the x-axis. 55 Figure 2-4. Kolmogorov-Smirnov two sample test for all relevés, comparing landslide and undisturbed plant abundance for Beatton River (BE), Cecil Lake (CE), and Hasler Flats (HA) study areas. Results where the maximum D statistic is greater than the critical value indicate a significant difference in plant community between two samples. 57 viii Figure 2-5. NMS (nonmetric multidimensional scaling) ordination biplot of 116 subjectively located averaged BEC (biogeoclimatic ecosystem classification) plots and 163 plant species sampled on three landslide study areas. The scale of the biplot has been increased to 700% due to the crowded distribution of the plots. Axis 1 explains 50% of the variation and represents an apparent gradient from forest community to open/grassland community, from left to right across the figure. Axis 2 explains 24% of the variation and represents an apparent soil moisture gradient from drier to wetter, moving from the upper to lower extent of the figure. Both axes are significant using the Monte Carlo test (p = 0.0040 and final stress = 18.20). Study areas are colour-coded, as shown in the legend. BE = Beatton River, CE = Cecil Lake, and HA = Hasler Flats. The environmental variables correlated with the ordination axes are: MesoslopePos (mesoslope position), Moisture (soil moisture regime), SlopeGrad (slope gradient in %), HLI (heat load index), and MatOrigin (material origin). Strength and nature of environmental variable relationships are represented by the length and direction of the red vectors. Undisturbed benchmark plots BE16-29, BE16-30, CE-35a, and HA16-01 are highlighted in yellow in the biplot. 59 Figure 2-6. Site series/types mapping example for Beatton River landslide study site. Site series/types codes (shown in blue) are 101 = Sw – Trailing raspberry – Step moss, 101$ = At – Rose – Creamy peavine, 103 = SwPl – Soopolallie – Fuzzy-spiked wildrye, 110$ = At – Highbush-cranberry – Oak fern, 111 = Sw – Currant - Horsetail, Gg = Grassland, Ws = Wetland/pond (swamp). Black lines represent site series polygon boundaries. The pink line is the perimeter of the undisturbed terrain study site surrounding the landslide. 64 Figure 2-7. Site series percent area paired graphs for Beatton River, Cecil Lake, and Hasler Flats study areas, with categories ordered from low (left) to high (right) moisture regime. 66 Figure 2-8. Site series Kolmogorov-Smirnov two sample tests for the paired study sites of each study area of Beatton River, Cecil Lake, Hasler Flats. For all three tests, the critical value is greater than the D statistic, indicating no significant difference. 70 Figure 2-9. Example of digitised geotyping on the Cecil Lake landslide. Geotype codes (shown in blue) are PO = Pond/wet area, RI = Ridge, RA = Raft, PI = Pillar, Fl = Flood deposits -seasonal, SW = Swale, GU = Gully. The black lines are polygon boundaries. 71 Figure 2-10. Rank abundance curves comparing geotype diversity on landslides and undisturbed terrain. 77 Figure 2-11. Relationships between relevé vegetation diversity and geotype diversity in terms of richness, Shannon diversity, Pielou’s evenness, and Simpson’s diversity for all three landslide study areas. Blue dots represent landslide sites and orange dots represent undisturbed sites for each study area. ix 80 Regression lines are not shown, as no significant regression relationship was found for any of the comparisons. Figure 3-1. Sampling transects with a series of 1 x 1 m quadrats along a measuring tape. Image on the left is Beatton River undisturbed transect BEtu1a. (Photo August 26, 2018) Image on the right is Beatton River landslide transect BEtl1a. (Photo July 18, 2017, by P. Burton. Used with permission.) 129 Figure 3-2. Landslide and undisturbed transect beta diversity (Bw) regression: vegetation vs environment. Blue symbols represent Cecil Lake transects, green symbols represent Beatton River transects, and orange symbols represent Hasler Flats transects. The blue dashed line is the regression line for landslide transect beta diversity (Bw). There was no significant regression for the undisturbed transects. 140 Figure 3-3. Cecil Lake landslide transect CEtl3c, showing high diversity of habitats as illustrated by Alnus spp. next to a pond, with sparsely vegetated southfacing slopes on the other side of the pond. (Photo August 25, 2018) 144 Figure 3-4. Cecil Lake landslide transect CEtl1, showing lower diversity of adjacent habitats as illustrated by expanse of level ground and relatively homogeneous vegetation. (Photo August 25, 2017) 144 Figure 3-5. Sweetclover (Melilotus spp.) carpeting the toe of the Beatton River landslide. Transect is BEtl4. (Photo August 16, 2017) 146 Figure 3-6. Vegetation diversity on the Hasler Flats landslide, illustrated by dense layers of varied forbs, short shrubs, and tall shrubs. Transect is HAtl1a. (Photo July 28, 2018) 147 Figure 3-7. Invasive/exotic plants Lactuca serriola (prickly lettuce), Sonchus spp., Melilotus spp., and Elymus repens (quackgrass) on the Beatton River landslide. Transect shown is BEtl1. (Photo August 17, 2017) 148 Figure 3-8. Example of moist cottonwood (Populus balsamifera ssp. balsamifera) habitat in undisturbed terrain at Beatton River. Transect is BEtu1a. (Photo August 26, 2018) 149 Figure 3-9. Relatively homogeneous vegetation (young aspen – Populus tremuloides) in undisturbed terrain at Hasler Flats. Transect is HAtu1a. (Photo August 10, 2018) 150 Figure 3-10. Beatton River landslide transect BEtl6, showing microsite diversity of hummocks, depressions, ponds, and fissures. (Photo taken September 5, 2018) 153 Figure 3-11. Hasler Flats undisturbed transect HAtu3 showing broken terrain. Transect traverses uneven terrain and small fissures for the first 15 m (foreground), then ascends a steep slope for 8 m (visible in the distance), and x 155 finally levels out (evident in the photo where the sky shows through). (Photo August 3, 2018) Figure 3-12. Cecil Lake undisturbed transect CEtu2 showing uniform, level terrain. (Photo August 17, 2018) 156 Figure 4-1. Landslide ponds study area – Mapsheet 94A (1:250,000) Peace River Region of northeastern British Columbia, Canada. The yellow areas indicate history of landslides along the Peace River and its tributaries, based on shape file linework by Severin (2004). Shape file linework imagery reproduced with permission. 169 Figure 4-2. Sample of landslide pond mapping. Location is Cecil Lake. Ponds are outlined in blue. Yellow lines indicate landslide features (previously mapped by Severin 2004). Severin landslide shape file lines used with permission. 172 Figure 4-3. Simplified anatomy of a landslide showing head, body, and toe. Adapted from Cruden and Varnes 1996. Credit: Transportation Research Board. 1996. Landslides: Investigations and Mitigation. Special Report 247. https://doi.org/10.17226/11057. Reproduced with permission from the National Academy of Sciences, Courtesy of the National Academies Press, Washington, D.C. 174 Figure 4-4. Frequency distribution graphs showing landslide pond size distribution in hectares (ha). The upper graph shows all landslide-generated ponds in Mapsheet 94A, while the lower graph shows the size distribution for landslide ponds ≤ 1.00 ha. 180 Figure 4-5. Total number of landslide ponds per geomorphic location. 181 Figure 4-6. Total landslide pond area (ha) by geomorphic location. 181 Figure 4-7. Mean pond size (ha) by geomorphic location on landslide. 182 Figure 4-8. Paired graph showing overall number of landslides per landslide type compared to number of pond-bearing landslides per landslide type. Landslide type names have been abbreviated to accommodate graph. Type abbreviations: CO = Compound; EF = Earth flow; MF = Mobile flow; MR = Multi-level rotational; RA = Ravelling; RE = Retrogressive rotational; RO = Rotational; SR = Shallow retrogressive. Note: No ponds were found on Ravelling (RA) or Earth flow (EF) landslide types. 183 Figure 4-9. Number of ponds per landslide type. 184 Figure 4-10. Pond area (ha) in each pond-bearing landslide type. 184 Figure 4-11. Mean pond size (ha) per landslide type. 185 xi Figure 4-12. Number of ponds on head per landslide type. 186 Figure 4-13. Pond area (ha) on head per landslide type. 187 Figure 4-14. Number of ponds on body per landslide type. 188 Figure 4-15. Total pond area (ha) on body per landslide type. 189 Figure 4-16. Number of ponds on toe per landslide type. 190 Figure 4-17. Pond area (ha) on toe per landslide type. 191 Figure 4-18. Very new pond on landslide (Beatton River Landslide). The most recent movement on the landslide occurred in 2015, so pond was approximately three years old at time of photograph. (Photo June 19, 2018) 192 Figure 4-19. Newer pond on landslide (Beatton River Landslide). Pond is on an older part of the landslide near the toe. (Photo August 9, 2017) 192 Figure 4-20. Persistent pond on landslide (Cecil Lake Landslide). Landslide occurred in 1998, thus the pond was approximately 20 years old at time of photograph. (Photo June 23, 2017) 193 Figure 4-21. Recent beaver gnawing activity on young sapling near pond (Cecil Lake Landslide). (Photo September 7, 2017) 199 Figure 4-22. Beaver pond on the Cecil Lake Landslide. (Photo June 22, 2017) 199 Figure 4-23. Older beaver gnawing activity on the Cecil Lake Landslide. (Photo September 10, 2017) 200 Figure 4-24. Recent beaver activity at the Hasler Flats landslide. Top image shows well-used beaver path (bottom centre of photo) leading from the sidescarp to a landslide pond. Lower image shows very recent felling of large aspen trees (Populus tremuloides) by beavers just above the sidescarp. (Photos August 14, 2018) 201 xii ACKNOWLEDGEMENTS Financial support for this study was provided by the Province of British Columbia Ministry of Forests, the BC Conservation Federation, Dr. Phil Burton’s NSERC (Natural Sciences and Engineering Research Council of Canada) funds, and NSTP (Northern Scientific Training Program) award funding. Further funding was made available through various UNBC student awards: UNBC research project award, UNBC travel grant, the Susan Stevenson Memorial Award, and the Al Nevison Award. I would first like to thank my supervisors Dr. Phil Burton and Dr. Marten Geertsema for agreeing to take me on as a student without firm funding in place and with only a kernel of an idea for a PhD topic. Thank you to Phil for all his advice and enthusiasm, for sharing his vast knowledge about disturbance ecology, and for always pushing me to reach further. Thank you to Marten for his steady support and encouragement, for somehow always finding funds to keep me employed, for opening the world of landslides up to me, and for providing me with interesting opportunities to get out into the field and learn firsthand. I also thank my other Committee members Dr. Paul Sanborn and Dr. Roger Wheate, and Richard Kabzems, MSc, PAg, RPF (Ret). I thank Roger for his direct approach and patience with my GIS questions. Thank you to Paul for his good nature, willingness to provide advice, and for sharing his wealth of knowledge on soils. And thank you to Richard for sharing his expansive expertise regarding deciduous and mixedwood ecosystems of the Peace River Region. I would not have been able to complete this work without the assistance of those willing to share their knowledge or provide a word of encouragement. Thank you to Ken Simonar for field training and mentoring on site-level ecological sampling and classification, and to Benita Kaytor for assistance with field data collection on Peace River landslides. Thank you to Kim Lutz for assistance with PCORD and for her all-around support. I also thank Dr. Darwyn Coxson for advice on PCORD. I appreciate the late Bruce Rogers for his willingness to answer all my questions about ecological classification over the years. His presence and contributions will be greatly missed. Thank you to Alexandre Bevington for help with my remote sensing and technological questions. Thank you to Phil Burton, Richard Kabzems, Benita Kaytor, Ken Simonar, Curtis Bjork, Nick Hamilton, and Marika Cameron for assistance with plant identification. Finally, thank you to my dear friends who believed in me and encouraged me all along, even when they didn’t understand the work I was doing. You know who you are. xiii DEDICATION I dedicate this PhD Dissertation to my Mom and Dad, who instilled in me a love of the natural world and the North, appreciation of simple things, a sense of humour regardless of life’s situations, and perseverance in the face of adversity. God Bless. ~In loving memory of my father, Richard Kress, who left this world June 2, 2019~ xiv Chapter 1. Introduction to landslide biophysical diversity research 1.1 Introduction Throughout the world, landslides are known for their destructive impacts on ecosystems, infrastructure, and human life (Geertsema et al. 2009). However, relatively little has been studied regarding the contribution of landslides to ecosystem health and diversity over various scales. Following a disturbance such as a landslide, succession is initiated, and new plant communities begin to form. Many factors influence the pathways of succession, including dispersal processes, substrate availability, nutrient and moisture levels, and competition (Clements 1916; Gleason 1917; Gleason 1926; Connell and Slatyer 1977; Pickett and White 1985). Landslides are unusual disturbances in that primary succession and secondary succession occur simultaneously, due to the presence of biological legacies (Walker and Shiels 2013). Successional processes can also be reset if the landslide reactivates. Landslides are thus challenging to study. This research aims to describe and compare biophysical diversity on landslides and in the surrounding relatively undisturbed terrain in the landslide-prone Peace River Region of northeastern British Columbia, Canada. The research is multidisciplinary, drawing on theories, applications, and literature of landslide ecology, geomorphology, plant ecology, remote sensing and spatial applications, ecosystem restoration and ecological engineering, and hydrology. This chapter will present an introduction to the research by first discussing the background and context, followed by the research problem, the research aims, objectives and questions, and the significance of the research work. The chapter ends with an outline of the structure of the dissertation. 1 1.2 Background to the study Globally, catastrophic landslides draw the attention of communities, governments, land managers, and researchers. Landslides can be large and destructive and often occur suddenly and rapidly (Cruden and Varnes 1996a). They can kill people and animals, destroy forests, and seriously damage buildings and transportation infrastructure. Landslides can also alter or obliterate habitat for fish and wildlife. The most common triggers of naturally occurring landslides are intense rainfall, rapid snowmelt, alterations in water level, volcanic eruptions, and earthquakes (Wieczorek 1996). Because of the impacts of landslides on nature and society, study of these events is ever-increasing. Broadly, landslides can be divided into prehistoric and historic types. Prehistoric landslides are those that occurred in the period before recorded history and they can be thousands of years old. Study of prehistoric landslides can help explain how and why a landscape formed as it did and can help predict the nature and frequency of future landslides over space and time. Prehistoric landslide evolution is analysed using signatures or clues left behind in the landslide, such as organic material that can be radiocarbon dated. Historic landslides are those that have occurred in the time since history was written down. It is usually easier to determine the cause and triggers of these landslides, as they have not been subjected to significant weathering or reworking. Historic landslides can also help predict future events, although perhaps on shorter time scales. The Peace River area features a unique combination of landforms due to various processes that created and changed the landscape over different scales of time. The valley and tributaries consist of thick layers of glaciolacustrine material from the former Glacial Lake Peace (Hickin et al. 2015). Over thousands of years, deep postglacial incision carved out the 2 current valley, creating steep slopes along the major river systems. When present on steep slopes, the glaciolacustrine material is unstable. The area has a long history of natural disturbance caused by landslides, interspersed with periods of stability. Some of the landslides were one-time events, while others appear to have reactivated, burying previous slides to some extent. Some landslides are large, such as the Attachie landslide that blocked the Peace River in 1976. There is evidence of many other landslides that are quite small “slumps”. Above the river valleys there are rolling hills and plateaus. Landslide ecology Landslide events are both erosional and depositional processes, removing existing natural features while at the same time creating new features on the landscape. These processes can result in a unique diversity of soil, vegetation, and microsite types across the landslide environment (Geertsema and Pojar 2007). Landslide ecology is a branch of landscape ecology that investigates how plant and animal communities respond and interact as a result of the changes caused by landslides. Some key components of study are nutrient cycling, soil development, plant adaptations, dispersal and colonisation, new mixes of native and nonnative species, and successional trajectories. Studies address these relationships on multiple scales of space and time. Landslide ecology also involves the integration of these biological learnings into management practices for slope hydrology, soil erosion, and slope stability (Walker and Shiels 2013). Geomorphology Geomorphology is the study of the physical or morphological features or properties (landforms) of the Earth’s outer crust in relation to geological features (Schaetzl and 3 Thompson 2015). Geomorphic or landform processes involve mechanical transport of organic and inorganic material (Swanson et al. 1988) and are strongly influenced by slope, pore water pressure, and soil cohesion. Landforms influence air and ground temperature, moisture availability, and nutrient availability at a site. They affect the flow of vegetative propagules, energy, organisms, and material through a landscape. Landforms also can affect the frequency and spatial distribution of other natural disturbances (e.g. fire) and control geomorphic processes that alter biotic features. Concepts of geomorphology are central to the present study, with its focus on geomorphic processes and recovery of landslides. Plant ecology Plant ecology is a subdiscipline of ecology that studies plant abundance and distribution, effects of environmental factors on plant abundance, and interactions among and between plants and other organisms (Keddy 2007). This discipline can be applied from the microsite to landscape scales, and from seasonal to millennial temporal scales. Plant ecology plays a central role in the present research, as the diversity of plant communities across the landslide is documented and compared to the surrounding area. The theories and principles of the discipline are incorporated as they relate to plant colonisation and abundance, successional trajectories, deterministic microsite features, and the influence of other organisms on plants. Remote sensing and spatial applications Remote sensing refers to any type of technology that captures an image from a distance. The most commonly used types of remote sensing are aerial photographs, satellite images, and more recently LiDAR (light detection and ranging). In this study a combination of LiDARconstructed DEMs (digital elevation models), drone imagery, LANDSAT (land-use satellite) 4 imagery, GIS (geographical information system) applications, and aerial photographs were used to identify, delineate, and analyse various components of biophysical diversity. Ecosystem restoration and ecological engineering Ecological or ecosystem restoration involves intentional activities by humans to start or accelerate recovery of an ecosystem to maintain or enhance ecosystem health. In many cases, the ecosystem has been damaged or destroyed by direct or indirect human activities. In other circumstances, the ecosystem has been degraded by a natural agent such as wildfire, flood, or landslide to the point where it is unable to recover to its historic successional trajectory. Restoration efforts attempt to regain this trajectory. The restored ecosystem will not necessarily recover its former state, developing instead along an altered trajectory (Burton 1991; SER Working Group 2004). In many industries, native plants are being used to help restore ecosystems damaged by both human-caused and natural activities (Polster 1997; Stokes et al. 2009; Walker et al. 2009). The selected plants help stabilise soils and slopes, control water flow, and increase wildlife habitat, among many other services. This practice of applying engineering principles to natural earth materials is referred to as geological engineering. The present study draws heavily on the concepts, theories, and findings related to ecological restoration and geological engineering when providing conclusions and recommendations. Hydrology Hydrology studies the pathways, distribution and storage, and quality of water, and includes the hydrologic cycle (Schaetzl and Thompson 2015). Water is very important in landslide processes. It can both trigger a landslide and transport debris and sediments downslope and 5 downstream. Soil strength properties are a function of soil water content (Meusburger and Alewell 2008), and landslide occurrence is closely linked to pore water pressure thresholds. Prolonged periods of rainfall often trigger landslides. The probability of slope failure is determined by the balance between precipitation and evapotranspiration by plants (Van Beek and Van Asch 2004), therefore vegetation plays a large role in this process. Mechanisms of landslide movement can significantly alter the hydrology of a slope, making it more unstable and changing soil development processes, which can also result in the formation of ponds and wetlands. Hydrological features such as gullies can limit the spatial extent of a slide (Geertsema et al. 2010). The present study incorporates hydrological principles and concepts when assessing landslide initiation and evolution. It similarly employs these theories to investigate persistence of water bodies formed by landslides. 1.3 The research problem a) Current state of research While the geological and geomorphological processes of landslides have been extensively studied throughout the world, much less research has focused on the ecological processes. In northern British Columbia, even the geological processes are not as well studied, and ecological processes have not been researched to any extent. This presents a challenge to landslide management, as it is important to understand the ecological processes of landslides to fully appreciate landslide evolution and recovery over multiple temporal and spatial scales. Some work has been done on plant succession and ecosystem recovery on landslides (Francescato et al. 2001; Dale and Adams 2003), with several studies occurring in tropical climates (Shiels and Walker 2003; Shiels 2006; Shiels et al. 2006; Restrepo et al. 2009). 6 These studies have helped build knowledge on topics such as the role of organic materials and birds in landslide recovery. In recent years, landslide ecology research has focused on the application of knowledge about landslide recovery processes to restoration practices (Turner et al. 1998; Pickett et al. 2009; Walker and del Moral 2009; Walker et al. 2009). These findings have contributed greatly to the advancement of a more holistic approach to landslide management. b) Literature gaps Although there are increasingly more studies on landslide ecology and recovery, peerreviewed literature quantifying biophysical diversity on landslides is lacking. Even less available is any research on landslide biophysical diversity in northern climates. In northern British Columbia (BC), there visually appears to be a wider diversity of plant communities and microtopography types on landslide surfaces as compared to the surrounding landscape (Geertsema and Pojar 2006). However, other than a coastal study (Smith 1986) there have not been efforts in the northern part of the province to quantify vegetative and environmental differences between landslides and surrounding terrain and analyse these differences to determine their significance. On a broader scale, studies incorporating this type of biophysical diversity analysis in other parts of the world do not appear to exist. The present study is the first known large-scale and comprehensive comparison of biophysical diversity on landslides and surrounding undisturbed terrain. c) Problem With changing climate, landslides in BC are projected to become more frequent and have greater magnitude, due to increased precipitation and degradation of permafrost (Geertsema 7 2006). It is expected there will be increased interest in prevention and mitigation of landslides, as well as restoration of ecosystems altered by a disturbance such as this. A lack of knowledge about succession, the plant assemblages that form, and the influence of biophysical factors may hinder the ability to manage landslides and carry out conservation measures. This research attempts to quantify these facets of landslide ecology to obtain a better understanding and provide lessons for restoration. 1.4 Research aims, objectives, and questions This research aims to quantify, describe, and analyse components of biophysical diversity on landslides in the Peace River Region of northeastern British Columbia. The research further aims to compare this vegetation and site diversity with surrounding, relatively undisturbed terrain where possible, and provide some recommendations for restoration and land management on landslides. Research objectives There are five main objectives of the study: (1) To quantify and analyse biophysical diversity on landslides and compare diversity with that found on the surrounding undisturbed terrain. (2) To quantify, analyse and compare the distribution of site-level ecological classification on landslides and on the surrounding undisturbed terrain. (3) To quantify, analyse and compare spatial turnover (beta diversity) of plant species and microsites on landslides and on the surrounding undisturbed terrain. 8 (4) To use the findings on biophysical diversity to assess whether geomorphic diversity is a predictor of vegetation diversity. (5) To quantify and analyse the presence and distribution of landslide ponds on an area of the Peace River Region. Research questions There are five key research questions to be answered in this study: (1) Are landslides demonstrably more biophysically diverse than undisturbed ecosystems? (2) To what extent do landslides rearrange the relative abundance of site-level ecological classifications on a slope compared to adjacent undisturbed terrain? (3) What is the extent of turnover of microsite and plant species diversity on landslides, and how does this compare to adjacent undisturbed terrain? (4) Is vegetation diversity on landslides significantly related to geomorphological diversity? (5) What is the distribution and abundance of landslide ponds at regional and local scales, and what are the ecological and management implications? A large portion of the research centres around three landslides in the Peace River Region, presented in Figures 1-1 (Beatton River), 1-2 (Cecil Lake), and 1-3 (Hasler Flats). These landslides are described in detail in Chapter 2. 9 Figure 1-1. Beatton River landslide. Top image is an oblique 3D presentation of an ortho mosaic draped over drone imagery (flown in 2016), at vertical exaggeration 1X. Bottom image is an aerial view of the landslide, illustrating temporary partial blockage of the Beatton River. (Aerial photo was taken October 20, 2015, by M. Geertsema. Used with permission.) 10 Figure 1-2. Cecil Lake landslide. Top image is an oblique 3D presentation of World Imagery layer draped over LiDAR imagery, vertical exaggeration 1X. Inset in top image is a photograph of the headscarp and part of the landslide, facing west. Inset photo was taken May 12, 2016. Bottom image is an ortho mosaic presentation of the landslide, showing variable topography and ponds. 11 Figure 1-3. Hasler Flats landslide. Top image is an ortho mosaic draped over drone imagery, vertical exaggeration 1X. Bottom image is an aerial view of the landslide, facing south. (Aerial photo on bottom was taken September 30, 2013, by M. Geertsema. Used with permission.) 12 1.5 Significance At a time when researchers are more frequently collaborating across disciplines or incorporating other disciplines into their research, the current study is truly a multidisciplinary endeavour. Overall, this research will contribute to the body of knowledge on disturbance ecology and restoration by describing and quantifying biophysical diversity of landslides in northeastern BC and providing recommendations for land management. Key contributions include measurement of alpha diversity and site-level ecological classification on landslides and comparison with nearby undisturbed areas, and quantification of geomorphological diversity on landslides with comparison to undisturbed terrain. An additional advance in knowledge is the quantification and analysis of spatial turnover (beta diversity) on landslides, which indicates the degree of differentiation among biological communities. Further contributions include quantification and description of surface water on landslides at a landscape level. All these learnings can enhance understanding of succession, recovery, and restoration of landslides and similar disturbances. The research findings may be especially useful in northeastern BC, where a new hydroelectric dam is under construction on the banks of the landslide-prone Peace River. 1.6 Structural outline of the dissertation In Chapter 1, the context of the research has been introduced and framed within the wider realms of scientific endeavours. The research questions and objectives have been outlined, and an argument for the value of this research has been presented. Finally, a structural outline of the dissertation has been provided. 13 In Chapter 2, a detailed background on alpha diversity will be presented and methods will be laid out regarding data collection and diversity analysis for vegetation BEC (biogeoclimatic ecosystem classification) plots, relevés, site-level ecological classification mapping, geomorphic type mapping, microtopography variation, and the multivariate ordination of vegetation data. A detailed description of results from several lines of analysis will be presented. The results will be discussed in the context of existing research literature. Conclusions and recommendations on the findings will be provided, followed by a list of references. In Chapter 3, background on beta diversity (i.e. turnover) will be provided, followed by a detailed presentation of the methods used for data collection and analysis for assessment of beta diversity on a series of field transects. Results will be presented, which will then be discussed in detail in the context of previous research and implications for management. Conclusions and recommendations on the findings will be provided, followed by a list of references. In Chapter 4, background on landslide ponds will be presented and then detailed methods will be laid out regarding data collection and analysis of landslide ponds over the area covered by geographical mapsheet 94A (Charlie Lake). The results will be presented, followed by a discussion of key findings. Conclusions and recommendations will finish the chapter, closing out with references cited. In Chapter 5, a summary of key findings and insights will be provided, followed by implications and recommendations for future applications. Finally, appendices will provide additional information to supplement the various chapters. 14 1.7 References Burton, P. 1991. Ecosystem restoration versus reclamation: the value of managing for biodiversity. Proceedings of the 15th Annual British Columbia Mine Reclamation Symposium in Kamloops, BC, 1991. The Technical and Research Committee on Reclamation. pp. 17-26. Clements, F. 1916. Plant succession: an analysis of the development of vegetation. Carnegie Institution of Washington publication 242. Connell, J., and R. Slatyer. 1977. Mechanisms of succession in natural communities and their role in community stability and organization. The American Naturalist, 111(982): 1119-1144. Cruden, D., and D. Varnes. 1996. Landslide types and processes. In: Turner, A., Schuster, R. (eds.), Special Report 247: Landslides Investigation and Mitigation. National Research Council, Transportation Research Board, Washington, DC, pp. 36-75. Dale, V., and W. Adams. 2003. Plant reestablishment 15 years after the debris avalanche at Mount St. Helens, Washington. The Science of the Total Environment, 313:101-113. Francescato, V., M. Scotton, D. Zarin, J. Innes, and D. Bryant. 2001. Fifty years of natural revegetation on a landslide in Franconia Notch, New Hampshire, U.S.A. Canadian Journal of Botany, 79: 1477-1485. Geertsema, M. 2006. Hydrogeomorphic hazards in northern British Columbia. National Geographical Studies 341. Utrecht. 184 p. Geertsema, M., L. Highland, and L. Vaugeouis. 2009. Environmental impacts of landslides. In Landslides – Disaster Risk Reduction. K. Sassa and P. Canuti (eds), Springer-Verlag. Berlin, Heidelberg. 650 pp Geertsema, M., and J. Pojar. 2007. Influence of landslides on biophysical diversity – A perspective from British Columbia. Geomorphology, 89: 55-69. Geertsema, M., J. Schwab, P. Jordan, T. Millard, and T. Rollerson. 2010. Ch. 8, Hillslope processes In Compendium of Forest Hydrology and Geomorphology in British Columbia, Volume2 of 2. Pike, R., T. Redding, R. Moore, R. Winkler and K. Bladon (eds). B.C. Ministry of Forests and Range, Forest Science Program, Victoria, B.C. and FORREX Forum for Research and Extension in Natural Resources, Kamloops, B.C. Land Management Handbook 66. Gleason, H. 1917. The structure and development of the plant association. Bulletin of the Torrey Botanical Club, 44(10): 463-481. Gleason, H. 1926. The individualistic concept of the plant association. Bulletin of the Torrey Botanical Club, 53(1): 7-26. 15 Hickin, A., O. Lian, V. Levson, and Y. Cui. 2015. Pattern and chronology of glacial Lake Peace shorelines and implications for isostacy and ice-sheet configuration in northeastern British Columbia, Canada. Boreas, 44(2): 288-304. DOI 10.1111/bor.12110 Keddy, P. 2007. Plants and Vegetation: Origins, Processes, Consequences. Cambridge: Cambridge University Press, First Edition. 706 p Meusburger, K., and C. Alewell. 2008. Impacts of anthropogenic and environmental factors on the occurrence of shallow landslides in an alpine catchment (Urseren Valley, Switzerland). Natural Hazards and Earth System Sciences, 8: 509-520. Pickett, S., M. Cadenasso, and S. Meiners. 2009. Ever since Clements: from succession to vegetation dynamics and understanding to intervention. Applied Vegetation Science, 12(1): 9-21. Pickett, S., and P. White. 1985. The Ecology of Natural Disturbance and Patch Dynamics. Academic Press. San Diego, CA. 472 pp. Polster, D. 1997. Restoration of landslides and unstable slopes: considerations for bioengineering in Interior locations. Proceedings of the 21st Annual British Columbia Mine Reclamation Symposium in Cranbrook, BC. The Technical and Research Committee on Reclamation. Restrepo, C., L. Walker, A. Shiels, R. Bussmann, L. Claessens, S. Fisch, P. Lozano, G. Negi, L. Paolini, G. Poveda, C. Ramos-Scharrón, M. Richter, and E. Velázquez. 2009. Landsliding and its multiscale influence on mountainscapes. BioScience, 59(8): 685-698. Schaetzl, R., and M. Thompson. 2015. Soils: genesis and geomorphology. Cambridge University Press, New York, NY. SER Working Group. 2004. The SER International Primer on Ecological Restoration.www.ser.org & Tucson: Society for Ecological Restoration International. Shiels, A. 2006. Leaf litter decomposition and substrate chemistry of early successional species on landslides in Puerto Rico. Biotropica, 38(3): 348-353. Shiels, A., and L. Walker. 2003. Bird perches increase forest seeds on Puerto Rican landslides. Restoration Ecology, 11(4): 457-465. Shiels, A., L. Walker, and D. Thompson. 2006. Organic matter inputs create variable resource patches on Puerto Rican landslides. Plant Ecology, 184: 223-236. Smith, R. 1986. Soils, vegetation, and forest growth on landslides and surrounding logged and old-growth areas on the Queen Charlotte Islands. BC Ministry of Forests Land Management report 41. 16 Stokes, A., C. Atger, A. Bengough, T. Fourcaud, and R. Sidle. 2009. Desirable plant root traits for protecting natural and engineered slopes against landslides. Plant Soil 324:1–30. Swanson, F., T. Kratz, N. Caine, and R. Woodmansee. 1988. Landform effects on ecosystem patterns and processes. BioScience 38(2): 92-98. Turner, M, W. Baker, C. Peterson, and R. Peet. 1998. Factors influencing succession: lessons from large, infrequent natural disturbances. Ecosystems, 1(6): 511-523. Van Beek, L., and T. Van Asch. 2004. Regional assessment of the effects of land-use change on landslide hazard by means of physically based modelling. Natural Hazards, 31: 289-304. Walker, L., and A. Shiels. 2013. Landslide Ecology. Cambridge University Press. New York, NY. 300 pp. Walker, L., and R. del Moral. 2009. Lessons from primary succession for restoration of severely damaged habitats. Applied Vegetation Science, 12(1): 55-67. Walker, L., E. Velázquez, and A. Shiels. 2009. Applying lessons from ecological succession to the restoration of landslides. Plant and Soil, 324(1-2): 157-168. Wieczorek, G.F. 1996. Landslides: investigation and mitigation. Chapter 4 Landslide triggering mechanisms. Transportation Research Board Special Report, (247). 17 Chapter 2. Biophysical alpha diversity of landslides and comparison with surrounding undisturbed terrain 2.1 Introduction and Background Landslides display a wide variety of sites, soils, and vegetation patterns compared to the surrounding undisturbed landscape (Geertsema et al. 2006) and they stand out visually from the adjacent terrain in terms of vegetation types and coverage, surface soils, and relief. Extremes in surface roughness, nutrients, and moisture are evident over very short distances on landslides. Landslides are both erosional and depositional (Cruden and Varnes 1996). The erosional process can remove deep layers of soil that have developed over centuries, effectively setting the clock back for pedogenesis (Geertsema and Pojar 2007). Landslides mainly change soil properties by exposing parent material, creating a variety of stages of pedogenic development (Geertsema et al. 2009). Where once a Brunisol existed, there now may be an Orthic Regosol. As material moves downslope, it becomes jumbled and turbated, resulting in unique soil layers and buried organic material (Phillips and Lorz 2008). The depositional process may transport large amounts of organic material from the slopes above, depositing it in one place near the toe of the slide. These organic components contribute an influx of nutrients in the depositional toe zone (Walker and Shiels 2013), and result in a loss of nutrients from the erosional scarp zone. Tree uprooting on the landslide can affect soil morphology, distribution of rock fragments, and evolution of regolith, at the same time both creating and enhancing layering and vertical contrasts (Phillips and Lorz 2008). Ultimately, erosion and deposition cause increases in extremes of nutrient regimes, facilitating a variety of plant community development trajectories. 18 The processes of erosion and deposition may also create many distinct geomorphic formations on the landslide. At the headscarp, erosion can result in steep, dry cliffs. Material moving at a high speed can gain momentum and form ridges and pillars on the landslide body as it comes to a stop (Geertsema et al. 2006). Subsequent erosion may create deep gullies, and additional material can be deposited in flows. Weathering transforms ridges and pillars over time, reducing surface roughness. In a rotational slide, the backward tilting at times creates a depression where a sag pond can form (Takaoka 2015). Spreads produce horsts and grabens (raised and lowered blocks, respectively) as the material pulls apart between faults, and water may accumulate in the grabens. Small ponds may also form anywhere there is a depression in the landslide surface. Overall, the topography formed by landsliding can result in extremes of moisture. Very dry sites often abut very wet sites, and very rich sites often lie close to very poor sites. In regions where much of the terrain is inherently unstable, landslide processes and the resulting variety of geomorphic landforms and site characteristics can create a very disturbed landscape in terms of vegetation processes, soil processes, hydrology, and habitat. Habitat diversity is strongly related to patterns of disturbance and recovery (Sousa 1984; Geertsema et al. 2009; Walker and Shiels 2013). If both site and soil change, the soil changes may persist much longer and have more profound ecological effects, such as the formation of wetlands or the infilling of valleys. Observations across northern British Columbia (BC), Canada, indicate landslides in this part of the province are much more biophysically diverse than nearby undisturbed terrain (Geertsema and Pojar 2007). However, biophysical diversity on landslides in northeastern BC has not been measured or quantified in any substantial way. This chapter investigates 19 whether landslides are demonstrably more biologically and physically diverse than adjacent undisturbed ecosystems. Four key questions were asked: 1) How does plant species abundance and distribution differ on landslides compared to adjacent undisturbed terrain? 2) To what extent do landslides rearrange the relative abundance of site-level ecological classifications on a slope compared to adjacent undisturbed terrain? 3) Is landslide geomorphology significantly more diverse than adjacent terrain, and 4) Is vegetation diversity on landslides significantly related to geomorphological diversity? This chapter assesses alpha diversity of vegetation and environmental sites (i.e. biophysical diversity) within landslides in the study area and compares it with the surrounding undisturbed terrain where possible. The chapter also investigates the possibility of correlations between vegetation diversity and geomorphic diversity. Biophysical diversity in this study refers to the variety of vegetation and physical sites; for vegetation this variety can include both compositional and structural elements (Pitkanen 2000). It is hypothesised that there is higher vegetation diversity, site-level ecological classification diversity, and geomorphic diversity on landslides compared to the surrounding relatively undisturbed terrain, and that geomorphic diversity positively influences vegetation diversity. Because of the complexities and many theoretical layers involved in the study of diversity, it is necessary to first examine the concept of biophysical diversity and its measurement to lay the groundwork for this research chapter. Alpha diversity In general, diversity in nature can be described in a series of scales forming a hierarchy. The three basic levels of diversity are alpha diversity, beta diversity, and gamma diversity 20 (Whittaker 1960). These three levels are related, and often, quantification of one level is required before calculation of another. Alpha diversity refers to within-site diversity, while gamma diversity is described at a regional level (resultant of alpha and beta diversities) and beta diversity is generally understood as change in diversity between sites. Alpha diversity addresses diversity at the stand or plot level and represents the range of species that may interact with each other (Noss 1990; Magurran 2004). Biological diversity or biodiversity represents the variability among organisms from all sources and is essentially a comparative science (Magurran 2004). At different scales and contexts, the biodiversity of plants describes the range of alleles or genotypes in a population, the diversity of species or growth forms in a community, or the diversity of vegetation types across a landscape (Noss 1990; Burton et al. 1992). Diversity can be partitioned into two components: element richness and evenness (Simpson 1949). Species richness refers to the number of species in a unit of study (McIntosh 1967). Evenness is the variability in species abundances: the more equal the proportion of abundances between species, the higher the evenness, and the more diverse the community (Magurran 2004). Additionally, abundance is a surrogate measure of niche size, and statistical models assume that abundance is in some way related to a species’ ecological importance (Magurran 2004). Alpha diversity describes richness and evenness of species of a particular stand or community or group of organisms. These concepts and measurements of diversity can be applied to various physical elements of ecosystems as well, including vegetation structure and substrates. 21 Measurement of diversity The measurement of species diversity is based on three assumptions: 1) all species are equal, 2) all individuals are equal, and 3) species abundance has been recorded using appropriate and comparable units (Peet 1974). Diversity statistics are classified as either species richness measures (McIntosh 1967), or heterogeneity measures which combine richness and evenness components (Good 1953). Evenness measures assess the departure of the observed species abundance pattern from the expected pattern in a hypothetical assemblage (Lloyd and Ghelardi 1964). Measurement of species richness In its simplest form, species richness is the total number of species in a sampling area. Species richness can be estimated from samples through species-area curves, parametric methods, and nonparametric estimators (Magurran 2004). Species-area curves are a wellknown measure and plot the cumulative number of species recorded as a function of sampling effort (Colwell and Coddington 1994). These curves can be extrapolated to give an estimate of total richness of the assemblage, and they illustrate the rate at which new species are found. Species richness can also be estimated using jack-knife statistics or bootstrapping, resampling methods that only require incidence data. Differences in species richness between samples can be assessed in various ways. Richness estimates can be compared by using richness estimators such as Chao1 (non-parametric estimator that takes into account rare species) or ACE (i.e. abundance-based coverage estimator) to deduce overall minimum estimates (Chao 1984). A more accurate method of assessment of differences is rarefaction, which uses the information provided by all species 22 collected to estimate the richness of a smaller sample. The two samples can then be compared directly. However, rarefaction is computationally taxing. It also assumes that individuals are randomly dispersed, which means that richness in clumped communities will be overestimated. Rarefaction curves converge at small sample sizes, so sampling size must be sufficient to characterise the community (Gotelli and Colwell 2001). Rarefaction did not appear suitable for the purposes of the present study due to the prevalence of clumped communities and relatively small sample size; therefore, it was not used. Measurement of species abundance/evenness Many different species abundance models have been devised to describe the relationship between the number of species and the number of individuals of each species. Models include the geometric series (Motomura 1932) and Fisher’s logarithmic series (Fisher et al. 1943). Some models are better than others for showing species abundance distributions, but none are equally applicable to all assemblages due to local variations and the dependence on local influencing factors (Magurran 2004). However, distributions generated by models can still provide insight into processes determining biodiversity, because of the linkage of abundance of species with successful competition for limited resources. One of the most common and useful methods of displaying abundance data is the rank/abundance plot. Species are plotted sequentially from most to least abundant on the horizontal (x) axis. Abundances are usually shown in log10 format on the y-axis to capture the full range of abundances of different species. Data sets can also be displayed as percentages or proportions to allow for comparison between samples (Whittaker 1965). The rank/abundance plot distinctly highlights contrasting patterns of species richness and 23 differences in evenness among assemblages and can be very useful for showing changes following a disturbance (e.g. Bazzaz 1975). The shape of the rank/abundance plot can be used to infer which species abundance model best describes the data. Diversity indices In addition to the various measures of species richness and species abundance, there are also several diversity indices that have been developed. A diversity index is a single statistic that incorporates information on both richness and evenness components and is essentially a measure of heterogeneity (Good 1953; Hurlburt 1971). The weighting that is assigned to one component relative to the other can markedly affect the level of diversity calculated and the way sites or assemblages are ranked (Magurran 2004). Because each diversity index emphasises either the richness or evenness component of diversity, there is no single perfect index. Species diversity indices can be used to compare communities, but different measures may produce different rankings of sites. There are both parametric and nonparametric measures of diversity. Parametric measures or indices depend on distribution of species abundances while nonparametric measures do not. One of the most well-known nonparametric diversity measures is the Shannon index or Shannon’s diversity index, H’ (Shannon 1948; Magurran 2004; Ortiz-Burgos 2016), which assumes that individuals are randomly sampled from an infinitely large community, and that all species are represented in the sample. The Shannon index estimates species diversity in a community by considering the number of species and their relative abundances (evenness). It represents the degree of uncertainty in predicting the species of a given individual selected at random from a community. The Shannon index H’ 24 usually ranges from 1.5 to 3.5 in temperate zones (Magurran 2004). The higher the H’ value, the higher the diversity in a particular community. Some disadvantages of the Shannon index are that it is constrained and usually yields low numbers (Magurran 2004), and it can be difficult to interpret since it confounds richness and evenness. Simpson’s index, D (Simpson 1949) provides an alternative method of describing diversity. It calculates the probability that any two individuals drawn at random from an infinitely large community will belong to the same species (or some other category). As Simpson’s index D decreases, diversity increases. Simpson’s index is heavily weighted towards the most abundant species in the sample, so is less sensitive to species richness (Magurran 2004). It is one of the most meaningful and robust diversity measures, as it captures the variance of the species abundance distribution. Simpson’s index is also much less sensitive to sample size than the Shannon index. A variation on Simpson’s index D is Simpson’s index of diversity, 1-D (used in this chapter), also known as the Gini-Simpson index, which represents the probability that two individuals randomly selected from a sample will belong to different species or categories. In this second formula, the greater the value of 1-D, the more diverse the sample. Both Simpson measures are on a scale of 0 to 1. Pielou’s evenness, J (Pielou 1966) measures the relative abundance of the different species making up the richness in an area (Magurran 2004). As evenness increases, so does diversity. A community where just a few species are dominant is less diverse than one where the abundances are more evenly distributed among several species. Evenness is reported on a scale of 0 to 1. 25 Hill (1973) devised a way of describing the relationship between indices by defining a diversity index as the reciprocal mean proportional abundances and classifying according to the weighting the indices give to rare species. Hill related this classification to the fact that diversity measures emphasise either species richness or dominance. The conclusion about whether one site is more diverse than another can thus depend on the choice of diversity measure. Hill numbers (qD) are a parametric class of true diversity measures that integrate species richness and species abundances and represent a hierarchy of diversity values. Essentially, Hill numbers are the ‘effective number of species’ or ‘species equivalents’ (MacArthur 1965, 1972), representing the number of equally abundant species that would be needed to give the same value of a diversity measure such as the Shannon index or Simpson’s index. Hill numbers present a simplified interpretation of results, since the units always denote the effective number of species, regardless of position in the hierarchy (Morris et al. 2014). The parameter q determines the sensitivity of the measure to the relative abundances and quantifies how much the measure discounts rare species (Chiu and Chao 2014). The main Hill numbers (q) are q = 0, q = 1, q = 2, and q = ∞. Hill number q = 0 is simply richness and. Hill q = 1 is the exponential of the Shannon index and corresponds to the weighted harmonic mean of the species’ proportional abundances. Hill q = 2 is the inverse of Simpson’s concentration index and is associated with the weighted arithmetic mean of abundances. Hill q = ∞ represents infinity. As q nears infinity, the weighted generalised mean with exponent q-1 approaches the maximum proportional abundance of the most abundant species in the set of data. The formula for Hill number calculations is provided in Section 2.3.2.3 of this chapter. 26 In ecology, diversity measures are normally calculated on data for living organisms, but they can also be applied to any set of measurable categories of items, such as mapped units of landscape features (Nagendra 2002; Ricotta and Avena 2003). These calculations are generally less computationally intensive than for species, but the results still provide a robust measurement of diversity. 2.2 Study areas The regional area of interest is the Peace River Region of northeastern BC, on landslides <100 years old in glaciolacustrine sediments. The area was subjected to advances of the Laurentide Ice Sheet on at least three separate occasions, with the most recent retreat occurring more than 27,400 years ago (Mathews 1978; Mathews 1980). Evidence of glacial events is presented in interglacial fluviatile gravel units. The study area is mostly within the Alberta Plateau of the Interior Plains Region subdivision of the Canadian physiographic classification system, and it is drained by the Peace River (Holland 1976). The Interior Plains are east of the Rocky Mountain Foothills and consist of plateaus, plains, prairies, and lowlands. The Plains are underlain by sedimentary rocks chiefly of Cretaceous age, primarily of the Fort St. John Group with thick series of shales and sandstones near the top. This area is also comprised of the Dunvegan Formation, which is hard cliff-forming sandstone, and the Smoky Group, which is interbedded shales and sandstones. A small portion of the study area is within the Rocky Mountain Foothills subdivision of the Eastern System of the Canadian Cordillera (Holland 1976). The Foothills are underlain completely by sedimentary rock, mainly from the Mesozoic age and consisting of a variety of limestones, siltstones, sandstones, and shales. The Foothills were covered by continental ice during the Pleistocene. 27 The regional study area is completely within the Boreal Forest region of Canada and the Boreal Plains ecoregion of BC (Demarchi 2011) and is also entirely within the Boreal White and Black Spruce moist warm (BWBSmw) biogeoclimatic unit, as defined using the Provincial BEC (Biogeoclimatic Ecosystem Classification) guidelines (DeLong et al. 2011). The climate of the area is continental, with low annual precipitation (Chilton 1981). Winters are cold and long, with frequent inputs of continental arctic air. Summers are warm and short but have long daylight hours that benefit agriculture. The most common soils in the area are Grey Luvisols, but Luvic Gleysols, Eutric Brunisols, Chernozems, Solods, Organic Soils, and Regosols are also present (Valentine 1978; Lord and Green 1986). Following a series of field reconnaissance inspections, three landslides and their surrounding undisturbed terrain were selected as smaller study areas within the regional area of interest: these are known as the Beatton River, Cecil Lake, and Hasler Flats Landslides (Figure 2-1). Each of these study areas consisted of two paired study sites: the landslide itself and a delineated equivalent portion of the immediate surrounding undisturbed terrain. All three landslides occurred near tributaries of the Peace River. The Peace River and its tributaries have a long history of extensive and recurring slope instability. In addition to the above criteria, the landslide study areas were chosen based on local knowledge, safe and efficient access, and the presence of surrounding relatively undisturbed terrain at least equal in area to the landslide. 28 Figure 2-1. Study areas of Beatton River, Cecil Lake, and Hasler Flats landslides, shown in Google Earth Pro. The landslides are in the Peace River Region of northeastern British Columbia, Canada. Landslide locations are outlined in yellow and identified with red locator balloons. Inset map shows landslide locations at a provincial scale and in relation to Mapsheet 94A, which is outlined in red. BE = Beatton River landslide, CE = Cecil Lake landslide, and HA = Hasler Flats landslide. The Beatton River landslide was the youngest disturbance, to a large degree a reactivation of older landslide deposits, with the most recent movement occurring in 2015. It is a rotational earth slide / earth flow and has somewhat more complex stratigraphy than the Cecil Lake or Hasler Flats slides, including exposed bedrock. The Beatton River landslide (56°21’57” N 120°42’26” W) is approximately 30 ha in size, with an elevation range of 655 m to 447 m (δ 29 208 m) and an average slope of ~17.2°. It is northeast facing, situated below (east of) a cultivated hay field and between mature forest stands to the north and a grassland/mature forest mix to the south. When the landslide occurred it temporarily blocked a portion of the channel of the Beatton River to the east. The Hasler Flats landslide is the smallest slide at approximately 1.5 ha. It is a southwestfacing spread which occurred in 2013 and is situated in post-glacial lake sand (4 m in thickness) over silty clay. It is below a young, regenerated deciduous cutblock to the east and between mature deciduous stands to the north and south. The Hasler Flats landslide (55°36’39” N 122°0’45” W) has an elevation range of 615 m to 598 m (δ 17 m) and an average slope of ~6.3°. It is adjacent to the Pine River to the west, and the slide deposited debris into the river channel when it first occurred. This small landslide was chosen due to its distinctive horst-graben formation and small size, two features which could be studied and contrasted against the other two landslides. It is also conveniently located along the highway west of Chetwynd and had easy access. The Cecil Lake landslide was both the oldest and the largest of the three selected landslides. It is a spread with a rupture surface in glacial lake sediments underlying glacial till and is largely comprised of silty clay. The Cecil Lake landslide (56°23’48” N 120°38’11” W) occurred in 1998 and is approximately 56 ha in size. It has an elevation range of 665 m to 556 m (δ 109 m) and an average slope of ~9.6°. It is a north-facing landslide less than a kilometre east of the Beatton River, and at its north boundary (i.e. toe) it contains a narrow but deeply incised unnamed creek that empties into the river. A small strip north of the creek was included in the landslide study site, as there was disturbance that appeared to be the same age as the rest of the landslide. The Cecil Lake landslide lies below a cultivated field and 30 mature deciduous and mixedwood forests at its south end where the headscarp is. To the east is an older landslide, while to the west the Cecil Lake landslide abuts mature mixedwood forests. To the north above the creek is a steep south-facing mosaic of grasslands and mixedwood forests. 2.3 Methods and Analysis 2.3.1 Subjectively placed BEC plots 2.3.1.1 Field sampling of subjectively located BEC plots In the first sampling season (2016), 91 subjectively placed 50 m2 plots were established on the three landslide study sites (Beatton River, Cecil Lake, Hasler Flats) using the Provincial BEC (Biogeoclimatic Ecosystems Classification) field sampling methodology (BC Gov 2010, 2015), modified to accommodate smaller plots. In addition, one 400 m2 (20 m x 20 m) benchmark mature forest plot each was established in the surrounding undisturbed terrain study site at Beatton River and Hasler Flats, along with one 50 m 2 benchmark grassland plot in the Beatton River undisturbed terrain study site. In 2017, an additional 21 50 m2 plots on the Cecil Lake landslide and one 400 m2 mature forest plot on the surrounding undisturbed terrain were sampled to complete coverage of the Cecil Lake study area, as seasonal limitations in 2016 ended the sampling prematurely. The objective of this sampling program was to assess and describe the variation in plant communities and site characteristics present on each landslide study site. The intent was to describe as much of the variability as possible through field reconnaissance across the landslide. In total 116 plots were sampled across the three study areas: 30 plots at Beatton River, 30 plots at Hasler Flats, and 56 plots at Cecil Lake. 31 Data were recorded on FS882 field cards normally used in British Columbia BEC field sampling (BC Gov 2010). Procedures followed the Describing Ecosystems field guide (BC Gov 2010, 2015) and provincial terrain classification guidelines (Howes and Kenk 1997). The Biogeoclimatic Ecosystem Classification (BEC) system is a hierarchical site-level ecological classification method developed in BC that combines climatic, vegetation, and site classifications (Pojar et al. 1987; Meidinger and Pojar 1991). The BEC system provides a framework for organising ecological information and ecosystem management learnings, creates common terminology among forest resource managers, and is used to prescribe and monitor treatments at the site level (DeLong et al. 2011). The biogeoclimatic subzone is the basic unit used in climatic classification and is a group of ecosystems that have developed under the influence of the same regional climate. Each subzone has a distinctive sequence of related ecosystems ranging from dry to wet sites, influenced not only by the regional climate, but also by local soil and topographic features. The site series is the basic unit of site classification and is defined by using late seral or climax vegetation. Site series represent site units with similar environmental properties and potential vegetation. An eight-class scale has been developed, based on relative soil moisture regime, relative soil nutrient regime, and various other environmental factors. The standard size of plot used in BEC sampling is 400 m2, or 20 m x 20 m. Sampling intensity and location of the subjective BEC plots was based on vegetative and geomorphic differences observed in the field, with plot locations determined while in the field traversing the study sites. Plots on the landslide were either circular or rectangular, depending on the extent and configuration of the geomorphic/vegetation type. Most plots were circular (i.e. 50 m2 – 3.99 m radius). If plots needed to be rectangular to capture a 32 microsite, they were configured to cover an area of 50 m2. The smaller plot size of the subjective plots was chosen due to the broken-up and small-scale nature of many microsites on the landslide. Plots in the surrounding undisturbed terrain were generally larger, as more extensive areas of distinct plant communities were present. At each plot, vegetation, soil, and environment information was recorded and representative photographs were taken. Plant species were identified and recorded using BC’s seven letter coding system (BC Forest Service 2016), and abundance was described in units of percent cover. Plant species identification was aided by consulting with specialists as well as several published and online resources (i.e. MacKinnon et al. 1992; Johnson et al. 1995; Kershaw et al. 1998; Douglas et al. 1998-2002; Klinkenberg 2021). For soils, information on geology, terrain, and organic and mineral horizons/layers was recorded. Environment information included microsite slope, microsite aspect, plant community structural stage, successional status, mesoslope position, surface substrate composition, topography, moisture regime, nutrient regime, and elevation. Soils were classified according to national standards (Soil Classification Working Group 1998). An initial attempt was made to classify each sample to BEC site series using the appropriate guidebooks, recognising that some sites are a complex of multiple site series, as is also often noted when mapping post-logging site types. Forested ecosystems or precursors to these types of ecosystems were classified using the BWBS zone field guidebook (DeLong et al. 2011). Wetland and water features were classified using the Wetlands of British Columbia guide (MacKenzie and Moran 2004) and Technical Report 68, Biogeoclimatic Ecosystem Classification of Non-Forested Ecosystems in British Columbia (MacKenzie 2012). Grasslands were also classified using the MacKenzie (2012) guide. Rationale for 33 classifications was supported by field photos and field notes, as well as analysis of and comparison with the digital imagery. For all classifications, it was recognised that landslide sites are seral or in a state of successional development as a precursor to supporting the mature plant community likely to develop on each site. The 2016 data were entered into the VPro [VENUS (Vegetation and Environment NexUS) PROfessional] database program (MacKenzie and Klassen 2004), which is used primarily by BC research ecologists and contains data fields that mirror the FS882 cards. The program allows transfer of entered data to a spreadsheet and generation of various reports for analysis. Data can also be directly exported for statistical analysis in ecological multivariate analysis programs such as PC-ORD (PC-ORD 2015). The 2017 Cecil Lake plot data were entered in an Excel spreadsheet, due to some technical problems with VPro. The subjectively located BEC 50 m 2 and 400 m2 plots were originally intended to gather general baseline information about plant and site characteristics of the study sites but were ultimately central in some analysis components of the research. Although most of these plots were smaller than the random 20 m x 20 m relevé main plots and were established using different criteria, and thus could not be used for direct comparison in some parts of the study, the plot information was instrumental in analysing plant and site diversity and mapping out site series/types. The subjectively located plot information was used to assist in mapping out polygons to describe site series/types and terrain diversity. Other key analyses using these plots were a nonmetric multidimensional scaling (NMS) ordination of the vegetation and selected environmental data and calculation of microtopographic variability. The plots could eventually be registered in the Provincial BEC database (BECWeb 2023) if the smaller size and variable configuration are acceptable. 34 2.3.1.2 Nonmetric multidimensional scaling (NMS) ordination of BEC plot plant species and environmental variables Nonmetric multidimensional scaling (NMS) ordination (Kruskal 1964; Mather 1976) was used to analyse vegetation and selected environmental data collected on the subjectively located BEC plots, to identify and describe any patterns in species composition in relation to environment. NMS is especially suited to heterogenous ecological datasets such as this one, because it does not require any specific design or model form, and so avoids parametric assumptions (Peck 2016). The multivariate analysis program PCORD v. 7 (McCune and Mefford 2016) was used to run the NMS ordination. Plant species and abundance data were loaded from a spreadsheet into the main matrix. Values for five select environmental variables were loaded into the secondary matrix. A random starting configuration was employed. The Gower distance measure was used, as some plots had no vegetation (as indicated by empty rows of data). The PC-ORD program user guide, citing previous works (Gower 1971; Legendre and Legendre 1998), notes that the Gower distance measure is a flexible, universal measure and is suitable for data sets containing empty rows, but not empty columns. Three runs with real data were carried out on the dataset. A Mantel test was performed to assess redundancy between each pair of runs. The dimensionality (i.e. appropriate number of axes) of the data was assessed by performing a stress test on autopilot mode at “thorough” setting and six axes, using 250 runs each of random and real data to determine the best solution. The stability of the final solution was assessed by checking to see if the stress leveled out and plateaued over time. The proportion of variance of each axis was calculated based on the r 2 (i.e. coefficient of determination) between distance in the ordination space and distance in the original species space. 35 The plant species most strongly associated with the environmental variables of interest were identified using Pearson correlations with the axes of the ordination. The five environmental variables assessed in the NMS ordination in the secondary matrix were soil moisture regime, mesoslope position, slope gradient, heat load index (HLI), and material origin. Heat load index and material origin variables were not specifically measured in the field but are derived variables, incorporating information from the field data. Heat load index is a measure of heat on a site based on slope, aspect, and latitude (McCune and Keon 2002). The index is derived from direct incident radiation, which is the radiant solar energy that hits the earth’s surface (Belessiotis and Delyannis 2011). To get a true representation of the heat potential for each plot, the aspect was “folded” about the northeastsouthwest line using the formula ABS(180-ABS(Aspect -225)) provided in McCune and Keon (2002). The following formula (Equation 3 in the 2002 paper) was then used to calculate heat load index for each plot: HLI = 0.339+0.808*COS(RADIANS(latitude))*COS(RADIANS(slope))0.196*SIN(RADIANS(latitude))*SIN(RADIANS(slope))-0.482*COS(RADIANS(folded aspect))*SIN(RADIANS(slope)) where latitude is the site’s location in degrees north of the equator, slope is in degrees, and folded aspect is calculated from aspect (in degrees azimuth from North) as shown above. The material origin variable was created to categorise the level of soil development on each plot. Five classes were identified, based on information obtained from the field data and photos. Class 1 was the most stable level of soil material development, representing mature in situ material in the surrounding terrain. Class 2 represented intact mature material on the 36 landslide that had been rafted from the surrounding terrain. Class 3 was material on the landslide body that had an intermediate level of soil development, with no A horizon and a weakly developed B horizon. Class 4 represented Orthic Regosols, having only a C horizon. Class 5 was ponds, with arrested development of soil, which in most cases was likely only a C horizon. Appendix 1 provides a more detailed description of each class. 2.3.2 Randomly located 400 m2 relevés 2.3.2.1 Field sampling of randomly located 400 m2 relevés In the 2017 and 2018 sampling seasons, three randomly placed 400 m2 (20 m x 20 m) square relevé plots were established on each landslide study site (Beatton River, Cecil Lake, Hasler Flats) to sample vegetation and environment components. Plot locations were determined beforehand using a grid system and random number generator. The same plot establishment and sampling procedures were followed for an equivalent area of the surrounding undisturbed landscape, for a total of six random relevés per study area, with three relevés on each of six paired study sites. In contrast to the subjectively located plots, the objective of this particular sampling program was to randomly sample vegetation and site conditions to compare and contrast mean diversity on the landslides and in the surrounding undisturbed terrain. Data collection methods for these plots were very similar to those used on the subjectively located 50 m2 and 400 m2 plots, following the same BEC field manuals cited in Section 2.3.1.1. For the sake of clarity and simplicity, however, the random plots will be henceforth referred to as relevés and the subjectively located plots will be referred to as BEC plots for the remainder of this paper. At each 20 m x 20 m relevé, vegetation, soil, and environment 37 details were recorded on FS882 field cards and representative photographs were taken. Environmental information was collected on slope, aspect, plant community structural stage, successional status, surface substrate composition, topography, mesoslope position, soil moisture regime, soil nutrient regime, and elevation. Vegetation cover was tallied by percent abundance and plant species were recorded using BC’s seven letter coding system (BC Forest Service 2016, 2020). Plant species were identified using the same resources as in Section 2.3.1.1. Individual tree species, diameters and heights were also recorded. Information was collected on geology, terrain, and organic and mineral horizons, but with a reduced focus on soils. Field data were entered into Excel spreadsheets. 2.3.2.2 Rank abundance curves of plant species in relevés Analysis of the relevé plot data was carried out in Excel. The mean relative abundance per species was calculated for the three relevés on each of the six study sites, and then the species were ordered from greatest to least percent cover. Mean rank abundance curves for the relevés were graphed for each study site, plotting the log base 10 of ranked plant cover abundance for each species. Individual curves were also plotted for each relevé. These rank abundance plots were examined and compared among study sites and between landslides and undisturbed terrain using the slope of a linear regression. The rank abundance method does not require a goodness of fit test, but rather equates diversity of the assemblage with the slope of the relationship, which reflects evenness of abundances. The steeper the slope, the lower the diversity of the sample, since the higher ranked species have much greater abundances than the lower ranked species. The richness of the sample is represented by the number of species on the horizontal axis. 38 2.3.2.3 Plant diversity indices and measures for relevés To assess alpha diversity on the landslide and compare within-slide alpha diversity with undisturbed terrain, plant species richness and various abundance measures were calculated for the relevé plot data of each landslide (n=3) and compared to diversity values obtained from the three relevés on the surrounding terrain. Species richness (S = total number of species), the Shannon index [H’ = -∑pi*ln(pi), where p is the proportional abundance of each species], Simpson’s index of diversity [1-D, or 1-∑(n/N)2, where n is the total abundance of a particular species and N is the total abundance of all species], and Pielou’s evenness value [J = H’/ln(S)], were calculated. To compare plant diversity between landslide and undisturbed sites for the relevés, two types of values were calculated. First, the overall diversity values were calculated by obtaining the mean cover value of each plant species over all three relevés for each study site. These means were then used to calculate richness, Shannon index of diversity, Pielou’s evenness, and Simpson’s index of diversity (1-D) for the study site. The second set of values was obtained by first calculating diversity values for each individual relevé in a study site, and then using these results to calculate the mean diversity values and the standard deviations for the study site. In addition, true diversity Hill numbers q = 0, 1, 2, and ∞ were calculated for the plant species composition of the relevés on each study site, using a pre-formulated Excel spreadsheet obtained online (Goepel 2012). The formula for the different Hill q values is: 39 Where: R = Richness q = Hill numbers order or effective number of species pi = proportional abundance of the ith species 2.3.2.4 Plant growth form and species abundance in relevés The composition and distribution of plant growth forms on a site can provide indicators of the environmental forces shaping an ecological community (Landau 2004). Abundances of individual plant species can also provide information about the influence of landslide disturbance on plant community development. To investigate a possible relationship between disturbance and growth form, each plant species in the three combined relevés on the landslide and the three relevés on the surrounding terrain for each landslide was classified and grouped by growth form in a table, along with each species’ average percent cover (abundance). The standard provincial growth form categories of tree, shrub, forb, graminoid, fern and fern allies, bryophyte, and lichen were used (BC Gov. 2010, 2015). The mean plant species abundances by study site were subtotalled for each growth form category and their proportions of the total cover were then calculated and compared among study sites. Each individual plant species was also ranked in descending order by mean percent cover for each study site and a list of the ten most abundant species by cover was compiled for each study site. Plant species autecology guidance resources were consulted for interpretation of results (Haeussler et al. 1990; FEIS 2023). 40 2.3.2.5 Two-sample Kolmogorov-Smirnov test to compare plant assemblages in relevés The two-way sample K-S test was used to test for significant differences between species abundance distributions of the two assemblages of relevés, for each of the three pairs of landslide-undisturbed terrain study sites. The two-sample Kolmogorov-Smirnov (K-S) test (Tokeshi 1993; Sokal and Rohlf 1995) is a nonparametric test of the equality of continuous or discontinuous one-dimensional probability distributions. It quantifies the distance between the empirical distribution functions of two samples and determines the likelihood of the two sets of samples occurring if they were drawn from the same (though unknown) probability distribution. The K-S test is considered one of the most useful nonparametric methods for comparing two samples, due to its sensitivity to differences in both location and shape of their empirical cumulative distribution functions. The maximum cumulative proportional difference (K-S test D statistic) of abundances between sites was calculated in Excel (Microsoft Corporation 2010) entering formulas by hand and compared with the critical value (1.36/√n), obtained from tables (Zaiontz 2017) based on a sample size >40 with a pvalue of 0.05. If the maximum D-statistic was greater than the critical value, the null hypothesis of no difference between samples was rejected. 2.3.3 Mapping, classifying, and analysing site series/types using multiple types of plot data 2.3.3.1 Site types/series The site type/series study incorporated the Beatton River, Cecil Lake, and Hasler Flats field data and photos from the BEC and relevé plots and a series of randomly located 30 m transects (described and analysed in Chapter 3) to type out vegetation and characterise patch 41 diversity on the landslides and the surrounding undisturbed terrain. In British Columbia, the “site series” is the key unit of site-level ecosystem classification categories, based on distinctive plant associations and typical soil, parent material, or slope position characteristics (described in Section 2.3.1.1). In this paper, categories will be referred to interchangeably as either site types or site series. Prior to mapping the site series, the perimeter of the landslide was first digitised on mosaic transparent drone imagery, and then an equivalent area of adjacent surrounding relatively undisturbed terrain was also digitised. This analysis was a combination of site series classification using field manuals and GIS mapping exercises in Global Mapper (Global Mapper 2020) using high-resolution drone (UAV – unmanned aerial vehicle) imagery (i.e. mosaic transparent group TIF (tag image format) file) and an underlay of LiDAR (light detection and ranging) DSM (digital surface model). Pre-processed, government-purchased LiDAR data was used, which was collected at an average of 1.1 to 1.3 points per m2. The data collection project was flown in 2006 at heights of 1200 to 1550 m using an Optech 3100 LiDAR system. In some areas the coverage was sparser, while in other places there was overlap and almost twice as many points per m2. The configuration of the outer perimeter of the undisturbed terrain was constrained by the limited availability of the same high-resolution imagery that was available for the landslide site series mapping. Prior to digitising site types/series polygons, an attempt was made to classify the site series of each relevé using the same manuals and guides as for the BEC plots (Section 2.3.1.1). When mapping the site types, plot data were employed to verify vegetation cover and serve as a means of ground-truthing. The digital drone imagery and field photos were used as crossreferences to confirm plot classifications. Although some preliminary site series classification was done on the BEC plots during the first field season (2016), these plot 42 classifications were reassessed based on a greater understanding of the local conditions following two more field seasons, as well as an assessment of the high-resolution digital imagery that subsequently became available. Classifications were revised where necessary. Site types/series were initially delineated and digitised on the digital imagery in Global Mapper and assigned classifications based on vegetation and identifiable environmental indicators such as aspect and steepness of slope. Aerial interpretation methods were applied to assist in distinguishing different types of vegetation (Sayn-Wittgenstein 1960; Avery 1969; Sayn-Wittgenstein 1978). The GPS (global positioning system) plot locations for all subjectively and randomly located plots for each landslide were then transferred into Global Mapper, complete with their site series classifications. Site series information determined from the plot data was used to verify the mapped types. Conversely, some plot site series classifications were modified after assessing the digital imagery and classification of other nearby plots. A plant indicator guide was consulted for site series that appeared transitional (Beaudry et al. 1999). The same steps for classification were followed for an equivalent area of the surrounding undisturbed terrain for each of the three study areas. Due to the transient and jumbled nature of vegetation, moisture, and soil material that is often characteristic of disturbed ecosystems such as landslides, not all plot sites fit into neat categories. As a result, some adjustments were made to the classifications. Additionally, some new site series/type categories were created to reflect site types influenced by human activities or types not described in the field guides, such as cultivated fields. 43 The areas of individual polygons of each site type were subtotalled, and proportions of each site series were calculated for the total area of each landslide and associated undisturbed area once the mapping was complete. The proportions of site series on and off the landslide were graphically compared in a two-way bar graph in order of moisture regime for each of the three study areas. Rank abundance curves were plotted for each study site, and KolmogorovSmirnov two-sample tests were done to compare landslide and undisturbed results and assess whether the two paired sites came from the same distribution or community. Finally, Shannon, Simpson’s (1-D), and Pielou’s diversity measures were calculated for the site types found at each study site and compared between landslide and undisturbed sites. 2.3.4 Mapping, classifying, and analysing biophysical features using multiple types of plot data 2.3.4.1 Geotyping To describe and assess geophysical diversity, distinct individual geomorphic features were mapped out on each landslide study site (Beatton River, Cecil Lake, and Hasler Flats). Each feature was classified as a specific geomorphic type using Global Mapper to view the imagery and digitise polygon boundaries. These features were referred to as geotypes for the purposes of this study. The same digital imagery and digitised perimeter boundaries used in the site series work described in Section 2.3.3.1 were utilised. The ultimate delineation for geotypes was based on terrain, although the presence of the “pond” geotype was at times first identified by the presence of cattail (Typha spp. – most likely Typha latifolia). The provincial terrain classification guidebook (Howes and Kenk 1998) was used as a baseline reference, and features were classified with one of the 44 categories from the guidebook where possible. However, classification into pre-existing categories was not always possible due to the nature of some landslide features on slides adjacent to fields or cutblocks. Geotype classification was assisted by reference to the field card data, notes, and photographs from the BEC plots and relevés, as well as from random transects sampled for the beta diversity chapter (see Chapter 3). The mapping was done on high-resolution drone photogrammetric imagery (mosaic transparent group TIF file), using Global Mapper. Digital elevation model (DEM) topographical imagery (DSM TIF file) was also used to assist in distinguishing geomorphic features. Once all features on each landslide were mapped and classified, the total areas of all the polygons for each geotype were added up. Summary statistics were then calculated. Further analysis of the geotyping results was carried out by calculating the Shannon and Simpson’s diversity indices, as well as Pielou’s evenness, for each landslide study site. Rank abundance curves were plotted for each of the six study sites to compare landslides and undisturbed terrain. Finally, a regression analysis was done to check for any relationship between relevé vegetation diversity and geotype diversity. 2.3.4.1 Microtopography/surface roughness of BEC plots Microtopography refers to the amount of soil surface roughness at the local level, at a scale that can fundamentally influence nutrient and groundwater regimes, and thus plant establishment and reproduction and wetland processes. Data preparation for assessment of microtopography on the study areas involved clipping out the buffered BEC plots from LiDAR or drone cloud points in Global Mapper and running an analysis on the standardised elevation of the points for each plot. The surface roughness/microtopography of each plot on 45 each study site was represented by the coefficient of variation (in %), calculated as the standard deviation of standardised point elevation values divided by the mean standardised elevation for each plot. Generally, the higher the coefficient of variation, the higher the spread of data relative to the mean standardised elevation, and thus the higher the microvariability of the terrain within the plot. Although the standard deviation of elevation method of assessing surface roughness used in this study is not as computationally intensive as some other methods, it has been shown to perform equally as well as other more complex measures (Rozycka et al. 2016). 2.4 Results 2.4.1 Diversity of relevés The landslide study areas exhibited a visually diverse array of plant communities and sites over short distances. Very dry, sparsely vegetated sites could be found juxtaposed with ponds and rafts, while level, heavily vegetated sites occurred next to steep, unvegetated scarps. An illustration of some of the diversity found in the study areas is presented in Figure 2-2. 46 Figure 2-2. Vegetation and site diversity on landslides in the study. From top left clockwise: horst and graben complex and mature rafts (Hasler Flats. Photo July 12, 2018); unvegetated weathered pillar next to heavy brush (Cecil Lake. Photo September 17, 2017); dewatered/revegetated pond site (Cecil Lake. Photo August 23, 2018); steep unvegetated scarps interspersed with relatively level swaths of abundant invasive vegetation and the occasional pond (Beatton River. Photo August 15, 2017). 47 Summaries of the ten overall most abundant plant species on landslide and undisturbed relevés at each study site are presented in Table 2-1. The species abundances are presented as mean percent cover of entire relevé area. A complete list of plant species and mean abundances for these plots is provided in Appendix 2. Table 2-1. Ten most abundant plant species for relevés at all study sites, comparing landslide and undisturbed results. Values are mean percent cover of the entire relevé area +/- standard error of the mean. 48 For the Beatton River landslide study site relevés, the forb yellow sweetclover (Melilotus officinalis) and the fern ally common horsetail (Equisetum arvense) were the leading species for percentage of total plant cover, at 28.8% and 12.2%, respectively. On the Beatton River undisturbed study site, the shrub saskatoon (Amelanchier alnifolia) and the tree paper birch (Betula papyrifera) were the leading species overall (10.8% and 10.7%, respectively), and wild sarsaparilla (Aralia nudicaulis) and twinflower (Linnaea borealis) were the leading forbs (7.8% and 6.8%, respectively). On the Cecil Lake landslide study site, the leading species overall was common horsetail at 33.5%. The leading shrubs on the Cecil Lake landslide study site were green alder (Alnus viridus ssp. sinuata) at 24%, followed by a willow (Salix sp.) at 5.6%. On the Cecil Lake undisturbed study site, the top two leading species were trees: white spruce (Picea glauca) at 22.9% and trembling aspen (Populus tremuloides) at 9.1%. The next most abundant species were the shrubs prickly rose (Rosa acicularis) at 6.9% and highbush cranberry (Viburnum edule) at 6.8%. For both Hasler Flats study sites, the fern ally common horsetail was the overall leading species (16.4% landslide, 15.6% undisturbed), followed by trembling aspen (10.8% landslide, 13.6% undisturbed). On the Hasler Flats landslide study site, seven out of ten leadings species were shrubs, while on the surrounding undisturbed study site, five out of ten leading species were shrubs. There was also a difference between landslides in the presence of undisturbed species that were also found on the landslide. For the Beatton River study area, none of the top ten undisturbed species were also found on the landslide. On the Cecil Lake landslide, four shrub species were shared between the undisturbed terrain and the landslide: Populus tremuloides, Rosa acicularis, Shepherdia canadensis, and Alnus viridis ssp. sinuata. Hasler Flats study area had six top species in common between the paired study sites: Equisetum 49 arvense, Populus tremuloides, Rosa acicularis, Alnus viridis ssp. sinuata, Cornus stolonifera, and Symphoricarpos albus. 2.4.1.1 Plant growth form composition and species abundances for relevés Plant growth form abundances varied significantly between relevé study sites. Table 2-2 provides a summary of mean vegetation covers by growth form as a percentage of mean total vegetation cover on the plots, as well as the subsequent proportions, for all six study sites. A complete list of abundances and proportions of all species by growth form in relation to mean total vegetation cover for each study site is provided in Appendix 3. 50 Table 2-2. Relevé mean growth form cover as a percentage of mean total vegetation cover, along with proportions, for all study sites. The table provides a comparison between landslide and undisturbed vegetation growth form cover. Study site Vegetation cover Mean total vegetation cover (% ) Beatton Landslide Beatton Cecil Undisturbed Landslide Cecil Hasler Undisturbed Landslide Hasler Undisturbed 59.68 97.35 104.16 87.02 81.69 100.81 Cover (%) 0.20 35.62 11.22 18.46 0.34 17.09 17.55 10.44 Proportion of mean total (%) 10.02 40.94 13.74 18.31 Cover (%) 1.74 37.27 43.83 27.57 41.53 46.04 Proportion of mean total (%) 2.92 38.29 42.08 31.68 50.84 45.67 Cover (%) 45.27 26.38 13.82 13.08 6.86 20.08 Proportion of mean total (%) 75.86 27.09 13.26 15.03 8.40 19.91 Cover (%) 0.13 15.97 1.67 2.75 2.97 0.39 Proportion of mean total (%) 0.21 16.40 1.60 3.16 3.63 0.39 Cover (%) 12.17 0.13 33.50 0.03 16.42 15.63 Proportion of mean total (%) 20.39 0.13 32.16 0.03 20.10 15.51 Cover (%) 0.17 0.43 0.90 7.93 2.69 0.21 Proportion of mean total (%) 0.28 0.44 0.86 9.11 3.30 0.21 Cover (%) N/A 0.08 0.02 0.05 N/A N/A Proportion of mean total (%) N/A 0.08 0.02 0.06 N/A N/A Trees Shrubs (% ) Forbs (% ) Graminoids Ferns & Fern allies Bryophytes Lichens (% ) For Beatton River, forbs dominated on the landslide (>75% of the total cover) while shrubs were leading on the undisturbed terrain (38.3% of total cover), followed by forbs (27.1%). For the Cecil Lake landslide study site, shrubs were dominant (43.8% of cover) followed by ferns and fern allies (33.5% of cover). On the Cecil Lake undisturbed terrain study site, trees dominated (41% of cover) followed closely by shrubs (>31% of cover). For the Hasler Flats landslide site, shrubs prevailed (>50% of cover) followed by ferns and fern allies (>20% of cover) and then trees (>13%). On the Hasler Flats undisturbed terrain, shrubs comprised 51 >45% of the total cover, followed by forbs (20% of cover) and then trees (>18% of cover). Fern and fern allies were close behind, at 15.5%. 2.4.1.2 Plant diversity indices and measures for relevés The results for relevé plant diversity assessment using both the mean cover values and the individual relevé cover values show that in all comparisons, the Shannon index and Pielou’s evenness were higher on the undisturbed terrain compared to the landslide (Table 2-3). However, all landslide study sites had more variation around the mean than their paired undisturbed study sites for the Shannon index. Table 2-3. Combined* and mean (+/- standard error) vegetation diversity indices for relevés Study site Beatton Landslide Beatton Undisturbed Cecil Landslide Cecil Undisturbed Hasler Landslide Hasler Undisturbed Combined Mean Shannon Shannon index index (H') (H') (n=3) Combined Pielou's evenness (J) Mean Pielou's evenness (J) (n=3) Combined Simpson's index (1-D) Mean Combined Simpson's Richness index (S) (1-D) (n=3) Me an Richness (S) (n=3) 1.75 1.78 ± 0.26 0.47 0.55 ± 0.10 0.72 0.59 ± 0.22 41 25.33 ± 4.04 3.22 2.58 ± 0.17 0.71 0.66 ± 0.05 0.95 0.89 ± 0.03 91 52.33 ± 13.01 2.46 2.25 ± 0.36 0.53 0.55 ± 0.08 0.84 0.82 ± 0.06 106 61.00 ± 18.25 2.65 2.43 ± 0.30 0.64 0.65 ± 0.08 0.90 0.88 ± 0.05 65 43.33 ± 4.04 2.82 2.64 ± 0.33 0.63 0.63 ± 0.07 0.92 0.90 ± 0.03 90 65.33 ± 4.93 2.94 2.63 ± 0.12 0.66 0.65 ± 0.01 0.93 0.90 ± 0.01 84 57.00 ± 10.15 *where combined values are based on first averaging plant abundance data from the three relevés and then calculating the diversity indices. Simpson’s index of diversity was higher on the Beatton River and Cecil Lake undisturbed study sites compared to the landslide study sites, with an especially marked difference between the Beatton River landslide and undisturbed sites. However, the Beatton River landslide had a much higher standard deviation than the undisturbed study site. Simpson’s 52 index of diversity was slightly higher on the Hasler Flats undisturbed site compared to the landslide site using the mean cover metric, while the two values were the same for the individual cover metric. Richness was lower on the Beatton River landslide compared to the undisturbed terrain, while it was higher on the landslide for both Cecil Lake and Hasler Flats. Hill numbers were calculated using the mean percent cover data of plant species of relevés for each study site, as shown in Table 2-4. Table 2-4. Hill numbers for vegetation diversity on relevés. Mean Hill Numbe rs -True Diversity qD: Generalised Beatton Beatton Cecil Cecil Hasler - Hasler Order q Mean Landslide Undisturbed Landslide Undisturbed Landslide Undisturbed 0 1 2 ∞ harm geom avg inf 25.00 6.04 4.20 2.63 52.33 13.36 8.85 4.38 61.00 9.84 5.61 3.23 43.33 11.74 8.01 4.39 65.33 14.54 9.68 5.15 57.00 13.94 8.86 4.37 For the Cecil Lake and Hasler Flats study areas, mean Hill number 0 (species richness) was higher on the landslide study site than on the undisturbed terrain study site, while at the Beatton River study area, Hill number 0 was much higher on the undisturbed terrain. For the rest of the Hill numbers 1, 2, and infinity (∞), the values were higher on the undisturbed terrain than on the landslide for the Beatton River and Cecil Lake study areas, but higher on the landslide than the undisturbed terrain study site for Hasler Flats. The Beatton River study area exhibited the biggest difference between landslide and undisturbed terrain study sites, for all four Hill numbers. The Beatton River landslide study site had the lowest Hill numbers of all three landslides while the Hasler Flats landslide study site had the highest Hill numbers. Overall, the Hasler Flats study area had the smallest differences in Hill number values between landslide and undisturbed terrain study sites, while the Beatton River study area generally had the largest differences. 53 2.4.1.3 Rank abundance curves for plant species on relevés Rank abundance curves provide a visual representation of the richness and evenness of a study site. The rank abundance curves for all relevé study sites are shown in Figure 2-3. Results indicate a higher evenness of the Cecil Lake and Hasler Flats undisturbed study sites compared to the landslide sites, as shown by the steeper gradient of the curves for the undisturbed sites. The opposite is apparent for the Beatton River study area, with the landslide study site having higher evenness than the undisturbed site. The rank abundance curve for the Cecil Lake study site highlights the greater richness on the landslide compared to the surrounding undisturbed terrain. The Cecil Lake landslide site also had the highest species richness overall. The Beatton River landslide study site had a much lower richness than the surrounding undisturbed site, with less than half the number of species (41 compared to 91). 54 Figure 2-3. Comparison of rank abundance distributions for relevé plot plant species for all study sites. Abundance rank by species is on the x-axis. 2.4.1.4 Kolmogorov-Smirnov two sample comparison for relevés The Kolmogorov-Smirnov two-sample test to compare the plant composition between the relevés on the paired study sites revealed that for the Beatton River and Cecil Lake study areas, the maximum proportional difference (D statistic) was greater than the critical value (Fig. 2-4). These results rejected the null hypothesis of no difference between the two samples, and indicated there was indeed a difference in plant community diversity structure 55 between the landslide and undisturbed study sites. The opposite was true for the two Hasler Flats study sites, where the maximum D statistic was less than the critical value by half. The null hypothesis of no difference could not be rejected. Overall, the Beatton River study area K-S two sample test had the biggest difference between maximum D statistic and critical value. 56 Figure 2-4. Kolmogorov-Smirnov two sample test for all relevés, comparing landslide and undisturbed plant abundance for Beatton River (BE), Cecil Lake (CE), and Hasler Flats (HA) study areas. Results where the maximum D statistic is greater than the critical value indicate a significant difference in plant community between two samples. 57 2.4.2 NMS ordination of BEC plot data: Assessing patterns and correlations of plant communities and environmental factors 2.4.2.1 Plant species and environment biplot of BEC plots Nonmetric multidimensional scaling (NMS) ordination using PC-ORD was carried out on the vegetation and environment data for all BEC plots (landslide and undisturbed benchmark) to seek pattern within a matrix of multiple responses, in this case plant species. The Mantel test for redundancy for the six pairs of runs of real data yielded values of 93.84%, 88.37%, 88.92%, 90.62%, 90.46%, and 86.00%, indicating high redundancy. The final (best) solution from the stress test had three dimensions or axes, with stress (i.e. residual sum of squares) of 18.20. Stress directly measures the quality of an ordination. The Monte Carlo test result used 250 randomised runs, with the probability that a similar final stress could have been obtained by chance being 0.0040, or significantly low. The final solution had 138 iterations, with the stress levelling out and plateauing, indicating stability of the solution. The resulting biplot (Figure 2-5) depicts all the plots in species space, paired with the environmental factors of interest. Axis 1 accounted for 50.7% of the variation in plant species composition, while Axis 2 accounted for 24.0% of the variation, totaling 74.7% of the plant community composition explained. Axis 3 is not shown, but accounted for 12.5% which, combined with Axis 1 and 2, explained a total of 87.2% of the variation in species composition among sites. The point distribution reflects the makeup of the plant community, with similar plots close together in the biplot. 58 Figure 2-5. NMS (nonmetric multidimensional scaling) ordination biplot of 116 subjectively located averaged BEC (biogeoclimatic ecosystem classification) plots and 163 plant species sampled on three landslide study areas. The scale of the biplot has been increased to 700% due to the crowded distribution of the plots. Axis 1 explains 50% of the variation and represents an apparent gradient from forest community to open/grassland community, from left to right across the figure. Axis 2 explains 24% of the variation and represents an apparent soil moisture gradient from drier to wetter, moving from the upper to lower extent of the figure. Both axes are significant using the Monte Carlo test (p = 0.0040 and final stress = 18.20). Study areas are colour-coded, as shown in the legend. BE = Beatton River, CE = Cecil Lake, and HA = Hasler Flats. The environmental variables correlated with the ordination axes are: MesoslopePos (mesoslope position), Moisture (soil moisture regime), SlopeGrad (slope gradient in %), HLI (heat load index), and MatOrigin (material origin). Strength and nature of environmental variable relationships are represented by the length and direction of the red vectors. Undisturbed benchmark plots BE16-29, BE16-30, CE-35a, and HA16-01 are highlighted in yellow in the biplot. The resulting statistically significant (randomisation p test = 0.0040) three-dimensional ordination solution shows the distribution of plots for all study areas (Beatton, Cecil, and 59 Hasler). Axis 1 and Axis 2 represent condensed gradients of differences in the composition of the data matrix. The environmental variables soil moisture regime, mesoslope position, heat load index, slope gradient, and material origin are represented as red vectors in the biplot, radiating out from the centroid. Axis 1 appeared to represent a gradient from forest community to open/grassland community, moving from left to right. The benchmark plots on the undisturbed terrain (highlighted in Figure 2-5 as BE16-29, BE16-30, CE-35a, and HA16-01) appeared quite different in composition from the associated landslides, as they were much further away from most plots in ordination space. These plots were also different from each other, based on their location on the biplot. Interpretation of Axis 2 is less clear, but it appears to represent a moisture gradient, transitioning from drier to wetter when moving from the lower to upper extent of the axis in ordination space. 2.4.2.2 Plant species correlations with axes using Pearson’s correlation coefficient r, for BEC plots Using the NMS ordination results, Pearson’s correlation r was calculated in PC-ORD for each plant species for Axis 1 and Axis 2. Plant species that were strongly correlated with Axis 1 either positively or negatively are shown in Table 2-5. The exotic forb yellow salsify (Tragopogon dubius) was most positively correlated with Axis 1, and the next four most positively correlated species were also grassland species. The forb creamy peavine (Lathyrus ochroleucus) was most negatively associated with Axis 1, followed by the shrub red honeysuckle (Lonicera dioica) and the tree trembling aspen (Populus tremuloides). 60 Table 2-5. Pearson correlations of BEC plot plant species abundance with NMS Axis 1. N = 116. Plant species that were most strongly correlated with Axis 2 are shown in Table 2-6. The strongest positive correlation was the shrub species red swamp currant (Ribes triste), followed by the forb red clover (Trifolium pratense) and the fern ally field horsetail (Equisetum arvense), and then the tree balsam poplar (Populus balsamifera ssp. balsamifera). The plant species with the strongest negative correlation was the forb western meadowrue (Thalictrum occidentale), followed by the shrub Sitka alder (Alnus viridis ssp. sinuata) and then an unknown fern species. Also showing strong negative correlations with Axis 2 were the forb large-leaved avens (Geum macrophyllum) and the pixie cup lichen (Cladonia pyxidata). Table 2-6. Pearson correlations of BEC plot plant species abundance with NMS Axis 2. N = 116. Positive correlation Plant species Common name Ribes triste Red swamp currant Trifolium pratense Red clover Equisetum arvense Field horsetail Populus balsamifera ssp. balsamifera Balsam poplar Fragaria virginiana Wild strawberry Negative correlation r 0.366 0.351 0.281 0.270 0.264 Plant species Common name Thalictrum occidentale Western meadowrue Alnus viridis ssp. sinuata Sitka alder Unknown fern sp. Fern sp. Geum macrophyllum Large-leaved avens Cladonia pyxidata Pixie cup lichen r -0.483 -0.400 -0.400 -0.392 -0.356 None of the environmental variables tested exhibited overly strong associations with either of the axes (Table 2-7). However, material origin (MatOrigin) showed a somewhat strong correlation with Axis 2, the apparent moisture gradient, while slope gradient (SlopeGrad) had a weak correlation with Axis 1, the forest-grassland gradient. Mesoslope position 61 (MesoslopePos), heat load index (HLI), and soil moisture regime (Moisture) all had weak correlations with Axis 1. Of the three variables associated with the forest-grassland gradient, mesoslope position had the strongest association, followed by moisture regime. Table 2-7. Pearson correlations of environment variables and Axis 1 and Axis 2 of the ordination space. N=116 Variable MatOrigin Moisture SlopeGrad MesoslopePos HLI Axis 1 (R2= 50.7) Axis 2 (R2= 24.0) 0.253 0.153 -0.023 -0.020 0.185 0.024 -0.086 -0.101 -0.103 0.010 PCORD does not report p-values for correlations between variables and ordination axes, because sample sizes are typically large enough that even a very small correlation is “statistically significant”. Therefore, usually the lowest r-value is more conservative than the one determined by the p-value (McCune and Grace 2002). 2.4.3 Mapping, classifying, and analysing site series/types using multiple types of plot data 2.4.3.1 Mapping, classifying, and analysing site types/series The site series digitising exercise yielded a variety of configurations on each of the study sites, with some very small site type polygons next to very large polygons. In total, 20 different site types/site series were identified (Table 2-8). 62 Table 2-8. Site types/series found on all landslides and undisturbed terrain in the study. Beatton Beatton Cecil Cecil Hasler Hasler landslide undisturbed landslide undisturbed landslide undisturbed "Site series" Description* 101 Sw – Trailing raspberry – Step moss 102 Pl – Kinnikinnick – Lingonberry 103 SwPl – Soopolallie – Fuzzy-spiked wildrye 110 Sw – Oak fern – Sarsaparilla 111 Sw – Currant - Horsetail 101$ At – Rose – Creamy peavine 103$ At – Rose – Fuzzy-spiked wildrye 110$ At – Highbush-cranberry – Oak fern 111$6B.1 Acb – Dogwood – Highbush-cranberry 111$6B.2 At – Cow-parsnip – Meadowrue 112 (Fm02) AcbSw – Mountain alder – Dogwood C Cultivated field Fl Flood deposits - seasonal FL Fluvial (creek/river) Gb Brushland Gb51 Saskatoon – Blue wildrye Gg Grassland Gg51 Sle nder whe atgrass – Pasture sage Ro Rock outcrop Rt Talus W Wetlands/ponds Totals m2 31190 977 79743 42487 30142 82840 0.0 8258 0.0 0.0 0.0 0.0 0.0 0.0 0.0 0.0 2009 0.0 242 23510 818 302216 m2 38200 0.0 35440 17430 57600 48945 13090 0.0 0.0 0.0 14460 0.0 0.0 0.0 0.0 46340 11740 17927 0.0 0.0 1044 302216 m2 161417 0.0 31387 30929 63888 134646 2257 38737 58348 0.0 0.0 0.0 6571 5989 0.0 0.0 0.0 0.0 0.0 0.0 27563 561732 m2 123924 0.0 0.0 6990 18784 250756 626 0.0 0.0 0.0 0.0 26670 0.0 4600 12550 0.0 62477 0.0 0.0 53537 1878 562792 m2 0.0 0.0 134 0.0 0.0 6097 1558 628 1257 1056 2770 0.0 0.0 0.0 0.0 0.0 0.0 0.0 0.0 0.0 1482 14982 m2 0.0 0.0 0.0 0.0 0.0 10130 0.0 807 886 0.0 3142 0.0 0.0 0.0 0.0 0.0 0.0 0.0 0.0 0.0 24 14989 *Sw=White spruce (Picea glauca ); Pl=Lodgepole pine (Pinus contorta ); At=Trembling aspen (Populus tremuloides ); Acb=Black cottonwood (Populus balsamifera ssp. balsamifera ) The majority of the site series in Table 2-8 are defined and described in the BWBS field guide (DeLong et al. 2011). As BWBS forests in the Peace River Region contain a notable component of mature deciduous trees, the field guide describes a number of deciduous site series, denoted by the dollar sign symbol $. There are four newly created or modified “site series” categories: cultivated field [C], flood deposits (seasonal) [Fl], fluvial (creek/river) [FL], and wetlands/ponds [W]. 63 Figure 2-6 illustrates an example of site series mapping on the Beatton River landslide. Figure 2-6. Site series/types mapping example for Beatton River landslide study site. Site series/types codes (shown in blue) are 101 = Sw – Trailing raspberry – Step moss, 101$ = At – Rose – Creamy peavine, 103 = SwPl – Soopolallie – Fuzzy-spiked wildrye, 110$ = At – Highbush-cranberry – Oak fern, 111 = Sw – Currant Horsetail, Gg = Grassland, Ws = Wetland/pond (swamp). Black lines represent site series polygon boundaries. The pink line is the perimeter of the undisturbed terrain study site surrounding the landslide. The results of the site series mapping and classification exercise showed a change in site series composition, area, and polygon count when comparing the landslide study site to the surrounding undisturbed terrain. The site series/types were graphed in order of moisture regime on paired bar charts, from driest to wettest, for each study area (Figure 2-7). For the 64 Beatton River landslide study site, the 101$ (i.e. mesic deciduous) site series occupied the highest proportion of the total study site area (27.4%), followed closely by 103, a somewhat drier site series, at 26.4 %. The next most abundant type on the Beatton River landslide was the moist site series 110 (14.1%), followed by the mesic coniferous 101 (10.3%) and then the very dry Rt site series (7.8%). The two mesic site series occupied 37.7% of the total landslide area. For the Beatton River undisturbed terrain study site, the leading site series was 111, a coniferous wet type (19.1%), followed closely by the deciduous mesic 101$ (16.2%) and the dry Gb51 (brushland) at 15.3%. The next highest abundance on the Beatton River undisturbed site was the coniferous mesic site series 101 (12.6%) and then the drier 103 (11.7%). Combined, the two mesic site series occupied 28.8% of the total undisturbed area. 65 Figure 2-7. Site series percent area paired graphs for Beatton River, Cecil Lake, and Hasler Flats study areas, with categories ordered from low (left) to high (right) moisture regime. 66 Overall, there were more site series polygons on the Beatton River landslide study site (43) than the undisturbed study site (29). The landslide study site had a higher percentage of mesic site series area, as well as a higher ratio of deciduous to coniferous mesic site series area. There was a similar number of mesic polygons on the Beatton River landslide and on the undisturbed terrain, but the mean sizes and ranges were markedly different. The landslide had a higher proportion of dry site series area compared to the undisturbed terrain. Although the landslide contained five times more pond polygons than the undisturbed study site, the undisturbed ponds were on average six times larger. For the Cecil Lake landslide study site, the mesic 101 site series per cent area was leading (28.7%) followed closely by its mesic deciduous equivalent 101$ (24.0%). The next two most abundant site series were 111 (11.4%) and 111$6B.1 (10.4%), which were wet types. For the Cecil Lake surrounding undisturbed terrain study site, the mesic deciduous 101$ site series area dominated (44.6%), followed by its coniferous equivalent 101 (22.0%). The next most abundant site series by per cent of total undisturbed area were grassland (Gg 11.1%) and talus (Rt 9.5%), which are dry types. Overall, mesic site series dominated on both the landslide and undisturbed terrain of the Cecil Lake study area, with coniferous 101 per cent area greater than deciduous 101$ on the landslide, and the opposite trend on the undisturbed terrain. There were more than four times as many site series/types polygons on the Cecil Lake landslide compared to the undisturbed terrain (381 polygons vs. 86), but almost 75% of the landslide polygons were ponds. There were 71 times more pond polygons on the landslide study site than on the surrounding undisturbed terrain. However, the ponds on the Cecil Lake undisturbed terrain were an average of almost five times larger than the landslide ponds. 67 On the Hasler Flats landslide study site, the 101$ deciduous site series was leading (40.7% of the total area), followed by a much wetter site series, 112 (Fm02), which was 18.5 %. The pattern was similar for the undisturbed terrain study site, with 101$ at 67.6 % and 112 (Fm06) at 21.0 % of the total study site area. The third most abundant site series on the Hasler Flats landslide was a very dry type, 103$ (10.4 %), followed by ponds (W) at 9.9 %, and then two very wet site series, 111$6B.1 (8.4 %) and 111$6B.2 (7.0 %). On the Hasler Flats undisturbed study site, the third most abundant site type was 111$6B.1 (5.9 %) followed by the moist site 110$ (5.4 %). Ponds occupied only 0.2 % of the total undisturbed area. Overall, the Hasler Flats landslide had more site series/types than the undisturbed terrain and there were significantly more individual polygons. The Hasler Flats landslide study site had eight site series, while the undisturbed terrain study site had five site series. The landslide had 47 polygons, while the undisturbed terrain had just seven polygons. The mean size of most polygons was smaller on the landslide. However, the pond polygons (W) on the landslide had a much larger mean size than those on the undisturbed study site, and there were also thirteen times more individual pond polygons on the landslide study site. The diversity indices of the site series for the study sites for all three study areas (Table 2-9) indicate that the Beatton River undisturbed terrain was more diverse in site series/type composition than the landslide for Shannon diversity H’, Pielou’s evenness J, and Simpson’s index of diversity 1-D. In contrast, the Cecil Lake and Hasler Flats landslide study sites were more diverse in site series /type composition than the surrounding undisturbed terrain regarding Shannon diversity, Pielou’s evenness, and Simpson’s index of diversity. In terms of richness, the value was the same both on and off the slide for Beatton River and Cecil 68 Lake study areas. The Hasler Flats study area, however, had a higher richness of site series/types on the landslide study site compared to the surrounding undisturbed terrain (8 site series vs. 5). Of the three study areas, Hasler Flats also had the biggest difference between each pair of index values comparing landslide and undisturbed study sites. Table 2-9. Diversity indices for site series polygons - all study sites. BE = Beatton River, CE = Cecil Lake, HA = Hasler Flats. Richness (S) Site BE -Landslide BE - Undisturbed CE - Landslide CE - Undisturbed HA - Landslide HA - Undisturbed 11 11 11 11 8 5 Shannon diversity index (H') 1.817 2.170 1.959 1.624 1.712 0.927 Pielou's Simpson's evenness index of diversity (J) (1-D) 0.758 0.808 0.905 0.872 0.817 0.823 0.677 0.728 0.823 0.766 0.576 0.493 The Kolmogorov-Smirnov two sample test for all three study areas (Fig. 2-8) showed that there was no significant difference between landslide and undisturbed site series diversity. This finding is evidenced by the fact that the critical value is higher than the maximum D statistic in all three cases. 69 Figure 2-8. Site series Kolmogorov-Smirnov two sample tests for the paired study sites of each study area of Beatton River, Cecil Lake, Hasler Flats. For all three tests, the critical value is greater than the D statistic, indicating no significant difference. 70 2.4.4 Mapping, classifying, and analysing biophysical features using multiple types of plot data 2.4.4.1 Geotyping: Mapping, classifying, and analysing geomorphic features Landslide Geotypes In the geotyping exercise, 20 different geomorphic types were identified and digitised over the three landslide study sites. A glossary describing each classification is found in Appendix 4. Figure 2-9 shows an example of the digitised geotyping on the Cecil Lake landslide study site. Figure 2-9. Example of digitised geotyping on the Cecil Lake landslide. Geotype codes (shown in blue) are PO = Pond/wet area, RI = Ridge, RA = Raft, PI = Pillar, Fl = Flood deposits -seasonal, SW = Swale, GU = Gully. The black lines are polygon boundaries. 71 The summaries of the resultant geotyped areas for each of the three landslides are presented in Table 2-10. Comprehensive tables listing each geotype polygon and its associated area for each landslide are provided in Appendix 5. For Beatton River and Cecil Lake, the slide matrix geotype occupied the largest percentage of the total area of the landslide study site. The slide matrix component was highest (almost 69%) on the Cecil Lake landslide and lowest (36.83%) on the Hasler Flats landslide. The leading geotype by percent area for Hasler Flats was rafts, at 45.72%. The second highest geotype percentage for both Beatton River and Cecil Lake was hummocks, at 14.25% and 10%, respectively, while for Hasler Flats the second highest geotype area was the landslide matrix (36.83%). The third and fourth highest geotype percentages were scarp (9.16%) and raft (8.54%) for Beatton River, raft (6.65%) and pond (4.91%) for Cecil Lake, and pond (9.89%) and scarp (6.03%) for Hasler Flats. 72 Table 2-10. Landslide geotype summary by area (m 2) and percentage. The number and composition of geotype polygons differed markedly among landslide study sites. Beatton River landslide study site had 56 separate geotype polygons, Cecil Lake had 327 polygons, and Hasler Flats had 39 polygons. For all three landslides, ponds had the highest geotype polygon count. On the Cecil Lake landslide study site, ponds comprised almost 87% of the total number of polygons, but they only added up to 4.9% of the total area. Ridges had the second highest polygon count on the Cecil Lake landslide, followed closely by rafts. On the Beatton River and Hasler Flats landslides, rafts had the second highest polygon count. The third highest polygon count for the Beatton River landslide study site 73 was represented by scarps, while for Hasler Flats scarps and ridges were tied for third place. The Beatton River study site had 15 different geotypes, Cecil Lake had 12 different geotypes, and Hasler Flats had five different geotypes. The diversity indices for geotypes (Table 2-11) showed that the Beatton River landslide had the highest geotype richness, Shannon index, and Simpson’s diversity of the three landslide study sites. The Beatton River landslide study site also had a higher Shannon diversity, Pielou’s evenness, and Simpson’s diversity than the oldest landslide, Cecil Lake. Hasler Flats had the lowest richness but the highest Pielou’s evenness. Table 2-11. Geotype diversity indices for landslide geotype polygons. Richness Shannon (S) diversity inde x (H') Site Beatton River Cecil Lake Hasler Flats 15 12 5 1.759 1.176 1.188 Pielou's evenness (J) Simpson's index of diversity (1-D) 0.650 0.473 0.738 0.739 0.506 0.642 Undisturbed Geotypes The results of the digitising of geotypes in the undisturbed terrain are presented in Table 212. Observations of the terrain in the field and on digital imagery showed a general trend of larger, more contiguous geotype polygons in the undisturbed areas for all three landslides. Digitising and summarising the polygons confirmed this, as all three undisturbed study sites had fewer types but larger, more extensive polygons. However, there appeared to be a greater diversity of geotypes on the landslides compared to the surrounding undisturbed terrain. 74 Table 2-12. Undisturbed geotype summary by area (m 2) and percentage. Diversity indices were calculated for the undisturbed geotypes (Table 2-13). Results show that the Cecil Lake study site had the highest richness, while Hasler Flats had the lowest richness. The Cecil Lake undisturbed terrain also had the highest Shannon index and Simpson’s index. The Hasler Flats undisturbed study site had the lowest Shannon and Simpson’s indices, but the highest Pielou’s evenness value. 75 Table 2-13. Geotype diversity indices for undisturbed geotype polygons. Richness Shannon Pielou's Simpson's (S) diversity index evenness index of (H') (J) diversity (1-D) Site Beatton River 8 1.547 0.744 0.743 Cecil Lake 9 1.599 0.728 0.761 Hasler Flats 6 1.450 0.810 0.731 Comparing landslide geotype diversity vs undisturbed geotype diversity Rank abundance curves for geotype data were plotted for each of the six study sites (Fig. 210). The results show that the undisturbed sites had a lower evenness overall, illustrated by the steeper curve. However, the landslide sites had higher richness for Beatton River and Cecil Lake. 76 Figure 2-10. Rank abundance curves comparing geotype diversity on landslides and undisturbed terrain. 77 2.4.4.2 Surface roughness/microtopography of BEC plots The results of the microtopography (surface roughness) exercise showed a range of coefficients of variation between the BEC plots for each landslide site, as shown in Table 210. The number of points for each buffered plot varied based on the quality of the LiDAR imagery and ranged from 48 to 120 points. A table summarising elevation point statistics for each buffered BEC plot is provided in Appendix 6. Table 2-14. BEC 50m2 plot LiDAR points: Elevation coefficient of variation (CV) summary statistics. Beatton River, which was the most recent landslide, had the highest mean elevation CV, whereas the Hasler Flats landslide had the lowest mean CV. The same pattern existed for the range of CV, where Beatton River landslide had the highest range and Hasler Flats landslide had the lowest range. Cecil Lake and Hasler Flats landslides had a much smaller minimum CV overall (22.46% and 29.93%, respectively) than Beatton River (41.82%). Beatton River also had the highest maximum CV, at 139.42%. All three landslides contained at least one plot with an elevation CV greater than 100%. Only a few plots were available for an assessment of the surface roughness on the surrounding undisturbed terrain for each study area, as the original plot sampling focused mainly on the landslide, with one plot in the undisturbed terrain for Cecil Lake and Hasler Flats and two plots in the undisturbed terrain for Beatton River. There were insufficient plots for a true comparison with the landslide plots. Additionally, the LiDAR point data used for analysing microtopography on the landslides study sites did not extend enough beyond the 78 landslide area to provide adequate coverage of the adjacent undisturbed sites to calculate elevation CV for these areas. However, observations obtained from traversing the undisturbed terrain during subsequent sampling for this chapter showed that this terrain was generally smoother and less varied than the landslide body. This was also evidenced by the greatly reduced number of ponds on the undisturbed study sites for all study areas. 2.4.4.3 Assessing correlation between vegetation diversity and geotype diversity Simple linear regression analyses were run to assess whether there was a significant association of vegetation diversity with geotype diversity in terms of richness, Shannon diversity, Pielou’s evenness, and Simpson’s diversity (Fig. 2-11). 79 Figure 2-11. Relationships between relevé vegetation diversity and geotype diversity in terms of richness, Shannon diversity, Pielou’s evenness, and Simpson’s diversity for all three landslide study areas. Blue dots represent landslide sites and orange dots represent undisturbed sites for each study area. Regression lines are not shown, as no significant regression relationship was found for any of the comparisons. 80 Regression results were obtained for landslide richness (R2 = 0.632, F (1,1) =1.718, p = 0.415), landslide Shannon diversity (R2 = 0.778, F (1,1) =3.495, p = 0.313), landslide Pielou’s evenness (R2 = 0.607, F (1,1) =1.546, p = 0.431), and landslide Simpson’s diversity (R2 =0.404, F (1,1) = 0.677, p = 0.562). In addition, regression results were obtained for undisturbed richness (R2 = 0.868, F (1,1) = 6.601, p = 0.236), undisturbed Shannon diversity (R2 = 0.798, F (1,1) = 3.944, p = 0.297), undisturbed Pielou’s evenness (R2 = 0.0376, F (1,1) = 0.0391, p = 0.876), and undisturbed Simpson’s diversity (R2 = 0.926, F (1,1) = 12.485, p = 0.176). For all comparisons, and contrary to expectations, geomorphological diversity did not significantly predict vegetation diversity. 2.5 Discussion This research chapter set out to answer some key questions related to the biophysical diversity of landslides in northeastern BC. The main hypothesis underpinning the research was that landslides are biophysically more diverse than nearby relatively undisturbed terrain, with a greater diversity of plant communities, species abundances, geomorphic types, and microsites. The methods employed to answer the questions consisted of a series of subjective and random vegetation sampling, as well as NMS ordination, species composition and abundance analyses, diversity index calculations, and classification and GIS mapping of biophysical features. The findings of this research point to a partial confirmation of the hypothesis that landslides are more diverse than the surrounding landscape, but this is not always the case. Results also indicate that landslide ecology is more complex than conventional succession theories would suggest. 81 1) How does plant species composition, abundance, and distribution differ on landslides compared to adjacent undisturbed terrain? Plant diversity of relevés Contrary to what was expected, the rank abundance curves and calculated evenness, Shannon, and Simpson diversity indices for the Beatton River and Cecil Lake study areas show the vegetation of the relevés on the undisturbed study sites is generally more diverse than on the landslides, while for the Hasler Flats study area there is no significant difference between disturbed and undisturbed relevés. The lower evenness on the landslides indicates a few species, especially exotics such as Melilotus spp. and Sonchus spp., are very abundant and there are many species with low relative abundances. These findings could be due to the surrounding terrain being much older than the landslide. As the landslides ranged between two and twenty years old at the time of sampling, not enough time had passed to allow the full spectrum of available plant species to establish. In addition, the headscarp was adjacent to an anthropogenically modified site for all three landslides: Beatton River and Cecil Lake landslides were both next to agricultural fields of forage crops, while Hasler Flats landslide was adjacent to a young deciduous cutblock. Anthropogenically developed or modified areas are usually relatively low in plant species diversity compared to the surrounding landscape. Further, the plants in cultivated agricultural fields are commonly grasses and other fast growing pioneer species which spread easily via wind and runoff. The greater richness on the landslide compared to the undisturbed terrain for Cecil Lake and Hasler Flats landslides in contrast to Beatton River may be because the Beatton River landslide study site was the most recently disturbed slide and had a higher proportion of bare 82 or sparsely vegetated areas. The Beatton landslide was largely a reactivation of a previous landslide. Thus, the randomly placed relevés were more likely to land on areas of low vegetation cover consisting of mainly exotic species. The Cecil Lake landslide study site had the highest species richness overall, likely because it was the oldest of the three slides and successional processes had allowed more species to establish. However, on the Cecil Lake landslide the leading species was still a fern ally (Equisetum arvense), while the second leading species was a shrub (Alnus viridis ssp. sinuata). Evenness measures how evenly species abundances are distributed in a plant community, and as its value increases, so does diversity. Mean Pielou’s evenness, J, was higher on the undisturbed terrain study sites compared to the landslide study sites for all three study areas, and all three undisturbed study sites had almost identical evenness values. In addition, the Beatton River and Cecil Lake study areas had identical evenness values on the landslide study sites, although Beatton River had slightly more variability. The paired Hasler Flats study sites had a much smaller difference between each other compared to Beatton River and Cecil Lake, and the Hasler Flats landslide had the highest evenness of the three landslide study sites. Overall, these findings suggest that the landslides are very similar in distribution of species abundances, despite the differences in sizes and ages. However, analysis of the top ten leading species on each study area revealed that although the three landslides all had high values of Equisetum arvense, Rubus idaeus, and Populus tremuloides, they each had a different array of leading species. The differences in species composition and abundances are most likely due to the propagule sources and amounts, as well as the available substrates on each landslide. 83 The Shannon index, H’, and Simpson’s index of diversity, 1-D, both indicated that vegetation was more diverse on the undisturbed terrain for Beatton River and Cecil Lake study areas but was very similar for the paired Hasler Flats study sites. Mean Shannon indices and mean Simpson’s indices of diversity were higher on the undisturbed terrain than on the landslide for the Beatton River and Cecil Lake study areas, but slightly lower on the undisturbed terrain compared to the landslide for the Hasler Flats study area. The Beatton River study area had the biggest difference in both Shannon and Simpson’s diversity between its paired study sites. The lower diversity on the Beatton River and Cecil Lake landslide study sites compared to the surrounding undisturbed terrain may be due to vegetation dynamics influenced by disturbance. Vegetation dynamics are driven by site availability, species availability, and species performance, and changes in any of these conditions can alter plant communities (Pickett et al. 2009). On the Beatton River landslide, Melilotus spp. and other exotics dominated on large portions of recently disturbed substrate, while on the Cecil Lake landslide, Alnus spp. and Equisetum arvense were widespread. Vegetation dynamics are intrinsically connected to landscape ecology, disturbance ecology, competition, invasion ecology, and community assembly. The process of recovery of landslide surfaces is complex due to the high spatial and temporal variability of soil stability and fertility (Walker et al. 2009). Surface soil erosion and patchiness of soil fertility can significantly hinder plant community development. The minimal difference in Shannon and Simpson’s diversity between the landslide and undisturbed terrain at Hasler Flats compared to the Beatton River and Cecil Lake study areas may be because the Hasler Flats landslide was much smaller, more gently sloped, and had a much high proportion of rafted material originating from the surrounding terrain, providing a 84 mosaic of stable and fertile substrates and propagules for new vegetation. Further, the adjacent cutblock above the headscarp at Hasler was approximately 15 years old, with established shrubs and forbs in the understory. A greater diversity of plant species was available from this source compared to above the headscarps at the Beatton River or Cecil Lake study areas. The biological legacies or residuals left after disturbance seem to be interacting with landslide size, age, and disturbance intensity in the study areas to influence vegetation dynamics, in agreement with other studies (Turner et al. 1998). Residual vegetation from vegetated rafts and chunks of intact soil can spread via seed banks, propagules, suckers, rhizomes, or serotinous cones. The life history traits of plants present at the time of disturbance interact with the disturbance intensity to influence the species composition of residuals. Succession occurs on a continuum of the role of residuals compared to new invaders, as well as a separate continuum of soil development (Franklin et al. 2000; Franklin and MacMahon 2000; Dale et al. 2005). Residuals are affected by the spatial variability and intensity of the disturbance, and thus larger disturbances such as the Cecil Lake landslide may present a greater degree of uncertainty and variability in successional pathways (Foster et al. 1998; Turner et al. 1998). Hill numbers represent true diversity, or the effective number of equally abundant species required to achieve a specific diversity measure value. For the Cecil Lake and Hasler Flats study areas, the Hill number q = 0 (richness) was higher on the landslide study sites, whereas on the Beatton River study area, q = 0 was higher on the undisturbed study site. However, for all other Hill numbers (q = 1, 2, and ∞), the value was higher on the undisturbed terrain for Cecil Lake and Beatton River but higher on the landslide for Hasler Flats. These findings 85 are consistent with the trends from the calculations of Shannon and Simpson’s diversities. The Hasler Flats study area has the smallest Hill number differences between paired study sites, while the Beatton River study area shows the biggest differences between paired study sites. These findings indicate once again that the Beatton River study area has the biggest difference in diversity between landslide and undisturbed terrain, and the Hasler Flats study area has the smallest difference. The lower diversity on the Beatton River landslide study site compared to the undisturbed terrain is most likely attributable to the young age of the landslide and the fact it is still quite active. In addition, the proximity of the hay field just above the scarp, along with the steep slope and prevailing winds, could have contributed to abundant reseeding with exotic species such as sweetclover (Melilotus spp.) as well as noxious weeds such as thistles (Sonchus spp., Cirsium spp.). As Hill numbers reached q = ∞, the differences on the undisturbed study sites became less different from each other, with values for all three sites being quite similar but all still higher than the landslide study sites. However, the landslide study sites maintained a distinct difference in value between each other, with the Hasler landslide study site having the highest Hill number value and the Beatton landslide study site having the lowest value. The maintenance of this difference indicates the sites are robustly different from each other in their diversities, and this difference appears to be influenced by spatial and temporal factors. Hasler Flats is the smallest landslide while Beatton River is the youngest landslide. The distribution of growth forms on a disturbed site can change over time and is an indicator of successional status (Dale and Adams 2003). The plant growth form abundances for the relevés differed significantly between paired study sites (landslide vs. undisturbed) and also between landslide study areas. On the Beatton River landslide study site, most of the cover 86 was forbs (>75% of total cover), followed by fern and fern allies (20% of total cover) and then distantly by shrubs (<3% of total cover). On the Beatton River undisturbed study site, shrubs were leading, followed by forbs and then trees and graminoids. The high dominance of forbs on the Beatton River landslide study site may be due to adjacency to a field of mixed forage species. In addition, the Beatton River landslide appears to be more active and is steeper than the other two slides, which could prevent more persistent woody species from establishing on the headscarp and large secondary scarps. The Beatton River landslide also had less pond cover and fewer wet areas in general, indicating a limited hydrological system. Although a large proportion of the forb species on the Beatton River landslide was exotic invasives, the landslide surface could still eventually become populated with persistent native species, as introduced species may switch from competition to facilitation in their relationship with native species. In a study of revegetation on a landslide caused by the eruption of Mount St. Helens, vegetation plots populated with nonnative species had higher vegetation cover and more native species richness than those sites not invaded, indicating any type of invasion may facilitate primary succession (Dale and Adams 2003). Annual plant species were most common in the first two years after the eruption, likely the result of wind dispersal and the ability to grow in poor soil. In that study, the introduced species were more successful due to their ability to establish, spread, and fix nitrogen. On the Beatton River study area, the prevalence of shrubs and forbs over trees on the surrounding undisturbed terrain is reflected in the dominance of the first two growth forms on the landslide. Although the landslide was surrounded mostly by mature mixedwood and deciduous forests, a significant portion of the landscape was a grassland/shrubland complex. 87 For both Hasler Flats and Cecil Lake landslide relevés, shrubs dominated, followed by ferns and fern allies. The dominance of shrubs on both landslide sites may be due to propagule contributions from rafts, as well as the advanced age of the Cecil Lake landslide, which could have allowed time for shrubs to seed in via wind and animal dispersal. On the undisturbed terrain, the leading growth form for Hasler Flats was again shrubs, while on the Cecil Lake undisturbed site trees dominated. The presence of more shrubs on the Hasler Flats undisturbed site may be the result of the deciduous, somewhat open overstory, while the Cecil Lake undisturbed terrain overstory was mainly coniferous or mixedwood with a denser canopy. As deciduous stands shed their leaves in the autumn, more sunlight and nutrients reach the understory, stimulating growth of shrubs in the spring. A comparison can be made of the autecology of the dominant species on the three different landslide surfaces. On the Beatton River landslide, the youngest slide, six out of the top ten most abundant species were exotics. Most of these exotics are perennials and can spread by both seeds and rhizomes. Wind and water are the primary modes of seed dispersal. The seeds were most likely transported downslope from the field above via prevailing winds and runoff. Most of the species are also seed-banking and have a longer range of seed viabilities or “shelf life” (FEIS 2023). The most abundant species on the Beatton River landslide, the exotic forb Melilotus officinalis (yellow sweetclover), dominated the toe of the landslide. It is an aggressive invader but is also a nitrogen-fixer, thus serving to improve the soil for future successional species. Melilotus officinalis is usually a biennial but is sometimes an annual or short-lived perennial and can have viable seed for up to 30 years (FEIS 2023). Thus, seed could be stored in the soil and reemerge following a future movement of the landslide. The four native species in the top ten most abundant list for the Beatton River 88 landslide are the fern ally common horsetail (Equisetum arvense), the forb mugwort (Artemisia sp.), the shrub red raspberry (Rubus idaeus), and the forb Lindley’s aster (Aster ciliolatus). All four species are perennials and can spread by both spores/seeds and rhizomes. At Hasler Flats, the second oldest and the smallest landslide, seven of the ten most abundant plant species were shrubs, and there was one tree species, one fern ally species, and one moss species. The five most abundant species were the fern ally common horsetail (Equisetum arvense), followed by the tree species trembling aspen (Populus tremuloides), and the shrubs prickly rose (Rosa acicularis), green alder (Alnus viridis ssp. sinuata), and red raspberry (Rubus idaeus). None of the top ten species were exotic. This difference in species composition and abundances compared to Beatton River can be explained by the presence of ample native seed sources in the form of multiple rafts down the slope of the slide, in a series of horsts and grabens. These rafts often spanned much of the width of the slide, providing many potential seed sources to populate exposed soil. In addition, because the slide was so small (<2 ha) and was abutted by mature forests on both flanks, seeds and propagules could be readily dispersed from the adjacent forest. On the Cecil Lake landslide, which was the oldest of the three slides and was 15 years older than the Hasler slide, five of the top ten most abundant species were shrubs, three species were trees, one species was a fern ally, and only one species was a forb. There were no exotic species in the list of the ten most abundant species. The five most abundant species were the fern ally Equisetum arvense, the shrub Alnus viridis ssp. sinuata, the forb palmate coltsfoot (Petasites frigidus), an unknown willow species (Salix sp.), and the tree balsam poplar (Populus balsamifera ssp. balsamifera). The common horsetail Equisetum arvense is 89 a moisture indicator. It absorbs silicon from the soil, and then in turn can absorb excess moisture. Equisetum arvense has a perennial rhizomatous stem system which can extend into the soil up to 1.8 m (FEIS 2023), and it also reproduces via spores. Alnus viridis ssp. sinuata also prefers moist sites. It disperses seed over long distances by wind or water and can fix nitrogen and stabilise slopes (Haeussler 1990). Petasites frigidus grows on moist to wet sites. It is a perennial forb that expands via a creeping root. Salix spp. in general prefer moister sites and produce large amounts of seeds which are dispersed by wind and water and germinate best on exposed mineral soil. Salix spp. also reproduce by sprouts, which can grow very rapidly. Populus balsamifera ssp. balsamifera is an indicator of high moisture and nutrients. Seeds are easily dispersed long distances by wind. Populus balsamifera ssp. balsamifera can regenerate from root suckers, stump sprouts, and buried branch pieces. Suckering is most abundant where mineral soil is exposed. The composition of the ten most abundant plant species on the Cecil Lake landslide is indicative of progression toward a more mature forested community. This makes sense, as the Cecil Lake landslide is the oldest of the three landslides. However, observations in the field showed large areas of the landslide at the north-facing headscarp that were still bare or sparsely vegetated. This delayed succession condition contrasts with a study of landslide recovery on a volcanic site (Saito et al. 2021), which found that grass vegetation recovery is quicker on shady slopes due to moisture and can recover within 12 years. It is likely the slower recovery of the north-facing headscarp at Cecil Lake is due to the colder climate of northeastern BC compared to Japan. In addition, there was evidence of reactivated mass movements along the Cecil Lake headscarp. 90 The two sample Kolmogorov-Smirnov (K-S) test showed the relevé communities of landslide vegetation were significantly different from the undisturbed plant communities for Beatton River and Cecil Lake study areas. These findings were consistent with the diversity values for the same relevés, which also indicated significant differences between the paired study sites for these two study areas. The K-S two-sample test also showed the two Beatton River study sites to be significantly more different from each other than the two Cecil Lake study sites were from each other. This pronounced difference on the Beatton River study area may be due to the younger age and active nature of the Beatton River landslide, in concert with the diversity of ecosystems (grassland, shrubland, mixedwood, mature aspen, mature conifer) in the surrounding undisturbed terrain. The K-S test did not show a significant difference in plant communities between the landslide and undisturbed study sites for Hasler Flats. This result also agreed with the diversity indices findings which showed that the plant communities were very similar on the landslide and on the undisturbed terrain. NMS ordination of plant and environment data from BEC plots In general, the NMS ordination of the plant and environment data from the BEC plots showed a combination clustered-scattered pattern, with few clear, strong associations. The concentration of more than half of each set of landslide plots in a central cluster surrounded by a somewhat uniformly scattered distribution of the remaining plots seems to indicate a significant degree of similarity among all three landslides. This redundancy may be due to the early successional stage of most of the plant communities, with different species still trying to establish and occupy a niche. 91 Although the ordination biplot did not reveal overly strong patterns, it did indicate an association of certain plant communities with landslide age. For the Beatton River landslide, the youngest slide, the plots that weren’t clustered at the centre were largely associated to the open/grassland side of the gradient. In contrast, the scattered Hasler Flats plots were mainly distributed on the left side of the biplot, indicating forested communities. The plant communities of the oldest landslide, Cecil Lake, appear to be much more diverse than on the other landslides, as the plot locations on the biplot that weren’t clustered at the centroid were scattered throughout much of the species space. For all three study areas, the plant communities of the benchmark plots in the undisturbed terrain appear markedly different from the landslide plots, as they all occur as outliers on the biplot. This increased distance in species space confirms field observations that the plant communities of the surrounding undisturbed terrain were distinctly different from the landslide communities. The benchmark plots are also quite different from each other, as they are all far apart on the ordination biplot. The Beatton River and Hasler Flats benchmark tree plots in undisturbed terrain are closer to each other on the biplot and thus more like each other in composition than either of the plots is to the Cecil Lake tree plot. This similarity between the two study sites is consistent with the field data, as the Beatton River and Hasler Flats plots were in deciduous stands, while the Cecil Lake plot was in a mixedwood stand. Of the environmental variables tested in the biplot, material origin and slope gradient appear to be bigger drivers of plant communities than mesoslope position, soil moisture regime, or heat load index. However, mesoslope position seems to have a greater influence on plant composition than soil moisture regime or heat load index. These findings are consistent with research on coastal BC comparing landslide revegetation with that of adjacent logged areas 92 of the same age (Smith 1986), where the main factors influencing the rate and pattern of plant recovery on the bare surfaces were associated rock types and slope position. On the landslides, the development of vegetation and forest productivity were influenced by the post-disturbance soil materials, which were modified by mixing, surface erosion, and sloughing. Smith (1986) found the revegetation rate was slower on the steep upper and middle thirds of a landslide compared to the lower slopes, where revegetation was quite rapid. He identified a gradient of plant communities along the slope gradient of the landslides. Conifers dominated on the middle and upper slide zones, especially on coarse and acidic materials. Red alder (Alnus rubra) was predominant on the lower parts of the slides, particularly in fine-textured materials. Shrub and bryophyte covers were higher on the logged areas, while the forb cover was the same on the landslide and the logged area. Landslides had a much lower productivity, producing only a third as much wood volume as the harvested areas after 60 years. In the Peace River Region study areas, field observations did not fully support the findings of Smith (1986) regarding the plant community gradient – slope gradient relationship. On the Hasler Flats landslide, shrubs and trees were interspersed down the length of the slope. On the Cecil Lake landslide, there were very few trees on the upper slope, scattered mixedwood rafts, young aspen (Populus tremuloides) patches and alder (Alnus spp.) swales on the middle slopes, and young (~10 years old) spruce (Picea glauca) and aspen scattered throughout the hummocky toe of the slide. The Beatton River landslide had very little tree cover overall, apart from some small rafts and a patch of Picea glauca seedlings in a rocky, rubbly portion of the slide. The main vegetation was the expanse of sweet clover (Melilotus spp.), thistle (Sonchus spp.), and other invaders occupying the hummocky toe of the slope. These 93 differences may be due to the younger ages of the landslides in this study, as well as the very different climates between the two studies. Positive Pearson’s correlations associated with the apparent successional plant community gradient (Axis 1) are strongest for exotic and grassland species and negative correlations are strongest with trees and shrubs. For Axis 2, the apparent moisture gradient, the species most positively associated were the shrub Ribes triste, followed by the forb Trifolium pratense, and then the moisture indicators Equisetum arvense (fern ally) and Populus balsamifera ssp. balsamifera (tree). 2) To what extent do landslides rearrange relative abundance of site series/types on a slope compared to adjacent undisturbed terrain? Site types/series Landslides appear to rearrange relative abundances of site types to varying degrees compared to the adjacent terrain, and this variation seems to be driven by the age of the landslide at least to some extent. Overall, the general trend for all three study areas was more site series/types on the landslide than in the surrounding undisturbed terrain, or in situations where there was the same number of site series, they differed in classification between the two paired study sites. Another general trend was a greater number of individual polygons on the landslide for each site series, even when the proportion of the site series was similar between slide and undisturbed sites. In the Beatton River study area, the landslide study site had a higher percentage of mesic site series and a higher ratio of deciduous to coniferous mesic sites compared to the surrounding undisturbed terrain. On the Beatton River undisturbed terrain study site mesic sites also comprised the largest proportion of the total 94 area. The Beatton River landslide and undisturbed study sites each contained a similar number of mesic polygons but the mean sizes and ranges were markedly different, and most mesic polygons were larger on the undisturbed terrain. The landslide also had a higher proportion of dry sites than the undisturbed terrain. Overall, there were more site series polygons on the Beatton River landslide compared to the undisturbed terrain. The slide had five times as many ponds, but the average pond on the undisturbed terrain was six times larger. Although there were some similarities with the other two study areas, the Cecil Lake study area had the most notable differences in site type distribution between landslide and undisturbed terrain among the three landslides. On the Cecil Lake study area, for both study sites mesic types dominated with deciduous 101$ greater than coniferous 101. In contrast, the next two most abundant site series/types on the landslide were wet types (111$6B.1 and 111), while on the undisturbed terrain the next two most abundant site series/types were very dry types (Gg -grassland and Rt -talus). The main reason for the difference of moisture gradient on the undisturbed terrain appears to be the influence of the south-facing slopes across the creek, whereas the Cecil Lake landslide itself is north-facing and seems to be heavily influenced by groundwater. The Cecil Lake landslide had more than four times as many site type/series polygons as the undisturbed terrain, but almost 75% of these were ponds. The average size of the ponds on the undisturbed terrain was almost five times greater than those on the landslide. On the slide, the ponds appeared in clusters just below the scarp and on the toe of the slope, where the terrain was more uneven and marked with depressions. The ponds in the undisturbed terrain were more stable and vegetated. Overall, the age, size, geology, and aspect may all have contributed to the distinctness of site type 95 distribution on the Cecil Lake landslide compared to Beatton River and Hasler Flats landslides. The Hasler Flats study area yielded site type distributions most like what might be expected when comparing landslides and undisturbed terrain. On the Hasler Flats study area, for both study sites the mesic 101$ was leading, followed by the much wetter 112(Fm02). However, on the undisturbed terrain 101$ was a much higher proportion than on the landslide. Further, on the landslide the third most abundant site series was the very dry 103$, while on the undisturbed terrain the third most abundant site series was the very wet 111$6B.1. These results indicate that on this smaller, younger landslide, mesic sites are reduced overall and there is more diversity of dry and wet site types compared to the undisturbed terrain. The landslide also has many more polygons and a few more site series than the undisturbed terrain. The mean size of the pond (W) polygons was much greater on the landslide, and there were 13 times more pond polygons than on the undisturbed area. The findings regarding larger average pond size on the landslide are opposite of the Beatton River and Cecil Lake study areas, but the findings about the greater number of ponds on the landslide compared to the undisturbed terrain were the same as for Beatton River and Cecil Lake. The larger pond size on the Hasler Flats landslide is likely due to the series of horsts (ridges) and grabens (trenches or depressions), with water accumulating in the grabens. The surrounding terrain is much more uniform, with fewer depressions containing persistent water. Overall, the contrasts in the Hasler Flats study area site type distributions are due to the movement type and subsequent features (horsts/grabens), and these differences are more pronounced because of the small size of the landslide. 96 The pond distribution patterns on the three landslides are consistent with a study in Japan (Takaoka 2015), which found that ponds mainly occur in displaced masses downslope from ridges and in linear depressions along main ridges. This pattern was most pronounced on the Cecil Lake landslide, which also had numerous small ponds at the base of the headscarp. The Japanese study also found that 90% of the ponds on the landslide were <1000 m2. This finding is consistent with the three landslides of the present study, where average pond size was 54.5 m2 (Beatton River), 97.1 m2 (Cecil Lake), and 56.9 m2 (Hasler Flats). The characteristic of small landslide pond size is also confirmed at a regional scale in the pond mapping exercise in Chapter 4 of this study. Although ponds on landslides occupy small areas, they are a very important component of biodiversity. They provide connectivity, shelter, and a water source for larger animals such as ungulates and birds. Pond size influences species richness, as a series of smaller ponds have more species and higher conservation value than a single large pond of the same total area (Oertli et al. 2002). Alpha diversity of site types on landslides appears to be influenced by the passage of time. Site series/type diversity was greater on the landslide than the undisturbed terrain for the Cecil Lake and Hasler Flats study areas in terms of Shannon diversity, Pielou’s evenness, and Simpson’s index of diversity. Richness was equal on the Cecil Lake paired study sites but higher on the Hasler Flats undisturbed terrain. Overall, the alpha diversity findings for Cecil Lake and Hasler Flats confirm observations in the field and indicate that site series are more diverse on older landslides compared to the surrounding undisturbed area. Conversely, site series diversity was greater on the undisturbed terrain for the Beatton River study area for all measures except richness, which was equal on the paired study sites. The findings for the Beatton River study area are consistent with observations in the field and indicate that site 97 series diversity is reduced on more recent, steep landslides which are still in a state of heightened activity. The lower site series diversity indices on the Beatton River landslide study site compared to the surrounding terrain and the other landslides are likely due to the landslide having large areas of uniform terrain and fewer rafts. The Hasler Flats study area shows the largest positive difference in site series diversity indices between the landslide and the undisturbed terrain. This difference may be because not only does the Hasler Flats landslide contain rafts and remnants of the surrounding more developed plant communities, but it also has ridges and depressions where new species have established, and new combinations of soil moisture and soil nutrient regimes are created. The small size of the Hasler Flats landslide also increases the proportions of those site series found on rafts. Overall, the differences in site series diversity and distributions between the Peace River Region study areas may be driven by both initial pre-slide vegetation and slope conditions and post-slide species interactions and geomorphic alterations over time. All three landslides in this study had evidence of past mass movements in the surrounding terrain, but the degree of diversity created by these past events differed between study areas. Vegetation may modify slope stability, thus influencing landslide size and severity, and at the landscape level vegetation may influence the spatial distribution of landslides through its interaction with the substrate (Restrepo et al. 2009). There may be both downslope and horizontal environmental gradients on a landslide, with conditions usually the mildest at landslide edges and harshest at the landslide centre. Over time, ecosystems on landslides can reorganise and embark on different successional trajectories, due to time lags caused by changing interactions, in addition to feedback between biota and substrate attributes. For example, differences in leaf 98 litter chemical makeup and rate of decomposition between some early successional species may influence organic matter dynamics and nutrient cycling rates (Shiels 2006). 3) Are landslides significantly more geomorphically diverse than the surrounding undisturbed terrain? Geotyping Geodiversity indices have been increasingly used to try to quantify geophysical diversity on larger areas as a proxy for species diversity (Wallis et al. 2021). The geodiversity index is the sum of the spatial diversity of multiple environmental variables measured within each site and its surroundings, and comprises climate, habitat, and soil. The index quantifies the degree of variation across space. However, there is mixed evidence as to how effective geodiversity is at predicting biodiversity and ecosystem functions at the regional scale, and Wallis et al. (2021) found that climate variables are more important predictors than habitat and soil variables at the regional scale. Overall, the results in that study showed the geodiversity index only explained a small amount of the variation in plants and ecosystems, while environmental conditions and resources explained most of the variation. While newer geodiversity indices may have limited utility, the geotype diversity work done in the present study with traditional diversity indices of Shannon and Simpson can be used for benchmark comparisons with plant and site series diversity. In general, the findings of the study highlight a relationship between geomorphic diversity and time, in view of the ages of the landslides. The geotyping analysis revealed that in terms of geomorphic diversity Cecil Lake, the oldest landslide, had the lowest Shannon diversity index, Pielou’s evenness, and Simpson’s index of diversity, while the Beatton River landslide yielded the highest 99 richness, Shannon diversity index and Simpson’s index of diversity. The Hasler Flats landslide registered the lowest richness but had higher Shannon diversity and Simpson’s index of diversity than the Cecil Lake landslide. The overall higher geomorphic diversity indices on the Beatton River landslide compared to the other landslides was likely because the slide was very recent, and there was still large-scale movement occurring. In addition, the Beatton River landslide was situated in more complex stratigraphy than either Cecil Lake or Hasler Flats, with exposed bedrock and boulder fields resulting from rockfall. The Beatton River landslide had some much steeper slopes, which were susceptible to slumping. Many of the geotypes on the Beatton River landslide—such as aprons, scarps, rubble, and blocky areas—were sparsely vegetated due to either their steepness or the lack of organic material. Sparse vegetation may also result from greater distances from a dispersal source. Studies have found revegetation occurs inward toward the slide body from the landslide edges and outward from islands or rafts of vegetation (Francescato et al. 2001). Some of the most sparsely vegetated geotypes at Beatton River were in the centre of the slide. The Cecil Lake landslide had the lowest geotype diversity indices of the three slides, except for richness. This seems contradictory at first, considering the variety of features observed in the field. However, features weather and become less distinct over time. The Cecil Lake landslide was very large, and some expansive areas were occupied by just one or two geotypes. Some examples of these geotypes include the long headscarp, as well as the large area of hummocks at the toe of the slide. The headscarp was quite sparsely vegetated, even though the landslide was twenty years old at time of sampling. The headscarp also still had mainly exotics and native grasses. This condition is likely due to the width and steepness of the slope, which can slow the rate of successful revegetation (Francescato et al. 2001). 100 Further, although most revegetation is initiated from the landslide edge, a significant length of the headscarp was bordered by a cultivated hay field. Therefore, the main source of plant propagules was likely from the field. Francescato et al. (2001) found that revegetation rate on landslides decreases over time, likely due to early saturation by easily established species. This situation may be occurring on the Cecil Lake landslide headscarp, due to the colonisation by agronomic species from the adjacent field. Hasler Flats and Cecil Lake landslides both contained ridges, with those on Cecil Lake much more prominent and extensive. The larger ridges on the Cecil Lake landslide were likely because the disturbance area was much bigger and steeper than the Hasler Flats landslide, so the material gained more momentum as it moved downslope. In addition, the material on the Cecil Lake landslide was silty clay, while on Hasler Flats the material was fine sand for the first four metres of thickness. The clay ridges would likely persist longer, while the sand ridges would not be as pronounced from the start, and they would become subdued with weathering. Although ponds did not occupy a notable amount of area on any of the slides, they were numerous and distributed throughout, especially on the Cecil Lake landslide. The trend of high numbers of small ponds was evident over a large area of the region, as described in Chapter 4. This abundance and widespread distribution of ponds indicated the presence of depressions throughout the slides, as well as the level of the groundwater. The ponds were in different stages of development depending on the slide and specific locations within the slide. Ponds provide habitat for many different types of wildlife. Evidence of wildlife use of the ponds was present, especially signs of beaver (Castor canadensis) activity such as felled trees, gnawed bark, and well-worn trails. 101 In all three study areas, vegetated rafts were a significant component of the landslide area. Rafts and other fragments of vegetative legacies are important for initiation of succession following disturbances. In one of only a few detailed quantitative studies of northern British Columbia landslides, Smith (1986) compared revegetation on landslides with surrounding logged areas as well as old growth areas on various islands of the northern coastal archipelago of Haida Gwaii. Initial revegetation on those landslides was shown to depend on the availability of stable microsites and islands of debris and other remnant organic material (Smith 1986). Microtopography/surface roughness It is assumed that over time, landslides will undergo smoothing of the surface due to slope degradation and local deposition of materials. The microtopography exercise in this study showed that all three landslides had a high degree of variability around the mean coefficient of variation (CV) of elevation on microsites and also a high mean CV overall, indicating variability of elevation is high both within sites and among sites. The most recent landslide, Beatton River, had the highest variability on average. This higher surface roughness could be due to the greater steepness of the slope in many places, as well as the presence of more rubbly areas and exposed bedrock and rockfall. It should be noted that only drone data were available for the Beatton River landslide for this exercise, so all elevation points were unclassified. Therefore, some of the points could be vegetation rather than ground points. However, there were only a few heavily vegetated plots, so the results are largely representative of ground surface elevations. The Beatton River landslide also had the highest range of elevational variability, which could be explained by somewhat equal proportions of steep, smooth slopes, rubbly moderate slopes, and hummocky gentle slopes at the toe. 102 The Cecil Lake landslide had the second highest elevational mean, maximum, and range of CV, even though it was older than the Hasler Flats landslide. This higher surface roughness could be mainly due to the steeper slope of the Cecil Lake landslide compared to the Hasler Flats landslide. Steeper slopes may increase the diversity of elevation variation as material falls and slides, creating rubbly piles at the same time as smooth scarps form. When compared with field photographs taken at the microsite level, individual plot elevational CV results were not always representative of on-the-ground plot characteristics. For example, on the Beatton landslide one of the highest CVs (Plot 1 at 68.58%) was a flat grassy raft. The high CV calculation may be due to the vegetation present, as bare ground points were not filtered out. Overall, however, the elevation CVs appeared to accurately reflect the plot characteristics. The results were also consistent with other microtopography research (Rozycka et al. 2016). 4) Is vegetation diversity on landslides significantly related to geomorphological diversity? One of the key premises of this study was that the diverse array of geomorphic features on landslides would positively influence the diversity of vegetation communities. Regression analyses were performed for each landslide to assess whether there were correlations between the relevé alpha diversity values of vegetation richness, Shannon index, evenness, and Simpson’s index and the same measures for geomorphic diversity, using the geotyping results obtained for both on the landslide and in the undisturbed terrain. All four regressions showed that there was not a significant relationship for any of the diversity indices tested in any of the three study areas. This was somewhat surprising, considering how field observations suggested that vegetation and site diversity were closely related. There was not 103 even a relatively stronger, but not significant, relationship on the landslides compared to the undisturbed terrain, as the R2 and p-values were similar for both study site categories. A possible explanation could be that more successional time is required before true relationships have become established and are statistically discernible. Alternatively, it could be that the scale of the exercise masked any relationships, as the geotyping was done at the landscape level. It is possible the sample size of three relevés per study site was simply too small to obtain an accurate result. It is also possible a different analysis would be more appropriate. More study is required on this aspect of the research. Limitations One of the limitations of the study design was the fact that study areas were partly chosen based on access. This naturally resulted in selecting landslides with road access and increased the likelihood that the slides would be next to a field or other managed landscape. The presence of development in turn likely influenced the establishment of exotics from the adjacent field. A more random selection of study areas could result in some more remote landslides where revegetation is primarily influenced by native forest vegetation from the surrounding terrain. Alternatively, a deliberate selection of some more remote landslides would have allowed for a focused or comparative study of natural succession in contrast with succession influenced by exotic species. In addition, it would have been beneficial to study more landslides, to better assess correlations between vegetation and geomorphic diversity. A further limitation was the small sample size of three landslides. Each landslide was unique in age, size, and slide type, which presented challenges when attempting to generalise findings. The differences among the landslides did provide a basis for identifying spatial and 104 temporal trends, which is very useful as a baseline dataset. However, a series of landslides from different age classes, size classes, and landslide types would help to complete the picture. The sample size for this study was selected based on time and resource constraints, but future studies could expand on the work presented here. An additional limitation of the study was the lack of sufficient LiDAR bare ground point data for assessing microtopography on undisturbed terrain. This deficit of imagery precluded a robust analysis of the undisturbed microtopography and did not allow a full comparison with the landslide study sites. The microtopography assessment component was added several years into the study and thus was not planned for, but for future landslide studies researchers should ensure adequate LiDAR is available before setting objectives and delineating study parameters where mapwork is planned. A related limitation was the limited availability of the LiDAR mosaic high-resolution imagery for the site series and geotype mapping on the undisturbed terrain for each study area. The outer perimeters were thus determined in part by the boundaries of the available high-resolution imagery. The timing of the field data collection posed another possible limitation. Due to funding availability and other scheduling factors, vegetation plot sampling was done from early summer into the fall. During the late summer and fall sampling, there was less foliage and it was thus more difficult to identify species and estimate cover. However, extra effort was made to identify species and estimate cover based on stems, fallen leaves, and plant crown expanses. Therefore, it is believed that species identification and cover estimates were accurate within an acceptable margin of error. Regardless, it is still preferable to conduct sampling when plants are at their most robust and easily identifiable during the growing season. 105 A further limitation of the study design was the restriction of landslides to the Peace River Region. The Peace River Region is somewhat unusual in the province in that most slides occur in glaciolacustrine material. Therefore, the results are more specific to this area and less generalisable. However, basic elements of succession and site variability can still be applied to other parts of the province, country, or world to see if the results are comparable. In addition, the findings could apply to any part of the world that has been affected by glaciation. Finally, a limitation of the data collection was the learning curve involved during some field sampling. This was especially the case when estimating plant species cover, particularly on the larger 400 m2 plots. The implications of this are that some of the initial plots may have slightly inaccurate cover estimates. An alternative to this approach could be to have more than one person estimating the cover on the first few plots, and then compare notes and calibrate measurements before proceeding, or to take drone imagery of each plot and assess plant species cover by image analysis. 2.6 Conclusions and recommendations Some general trends appear in the analysis of biophysical alpha diversity on landslides in the Peace River region of northeastern BC. Overall, plant communities vary depending on the age and size of the landslide and the slope and soil development of the various geotypes present. Exotic forb species are more dominant in the early stages of a landslide’s successional development. Shrubs and trees become more established as the landslide stabilises. However, landslides can once again reactivate, setting plant succession back on some areas of the landslide. 106 The landslides in the study tend to have more diverse site series units compared to an equal area of the surrounding undisturbed terrain. In general, the landslides contain a larger proportion of mesic sites compared to surrounding undisturbed terrain, but they also have more pronounced extremes of site series at either end of the soil moisture regime gradient. The site series polygons are generally smaller on the landslide, compared to larger swaths of intact site series in the undisturbed areas. The landslides in the study area are more geomorphically diverse than the surrounding undisturbed terrain. Geomorphic types tend to be more diverse on the landslides, due to mass movement, rearrangement of stratigraphy, and erosion of unstable substrates. Geomorphic diversity also varies inversely with the age of the landslide. Surface roughness also appears more diverse on landslides and tends to decline with landslide age. The most recent landslide had the highest surface roughness. Ultimately, landslides in the study are generally less diverse than surrounding undisturbed terrain regarding plant diversity, but more diverse in abundance and distribution of site series and geomorphology. Although the landslides are lower in plant diversity due to rapid and wide-reaching establishment by invasive species and persistence of early successional species, the site series and geomorphological diversity present provide conditions for a greater variety of plant communities and wildlife habitats over time. The findings of this study of alpha diversity on landslides in northeastern BC provide the groundwork for several recommendations regarding landslide restoration, management, and research. Landslides targeted for restoration should be prioritised and selected based on the risk of future instability to human safety and health, infrastructure, and ecosystems. 107 Remotely located landslides may be left to revegetate and restabilise unaided if future movements do not threaten to block waterways or cause heavy sedimentation. Restoration should first focus on landslides near communities and infrastructure, such as Old Fort and the Site C hydroelectric dam along the Peace River. Restoration should also focus on landslides near cultivated areas, as well as next to rivers, creeks, or other water bodies used for human drinking water or designated as sensitive wildlife habitat. Restoration of landslides is essentially managed succession (Walker and del Moral 2008). Long-term research on primary succession on landslides provides information on temporal vegetation dynamics that serve as lessons for restoration practices. Applicable learnings include information on site amendments, development of community structure, nutrient dynamics, species life history traits, species interactions (competition), and modeling of transitions and trajectories. When considering restoration of landslides, it is important to realise that no plant community is ever static (Pickett et al. 2009). Successful restoration depends on accurate identification of where on the temporal gradient of succession a site is located, and appropriate intervention in disturbance, colonisation, and competition processes. Landslides typically cause a loss of vertical vegetative structure, soil nutrients, and soil seed bank (Shiels and Walker 2003), as was observed on the three landslides studied. Restoration on landslides is particularly challenging due to the high spatial and temporal heterogeneity of soil stability and fertility (Walker et al. 2009). Surface soil erosion is a serious problem, as is patchiness of soil fertility. These conditions were especially evident on the Beatton River landslide but were also widespread along the headscarp of the Cecil Lake landslide. Persistent soil erosion is the first barrier to succession and restoration on landslides, and 108 measures to reshape slopes are often unsuccessful because of subsequent erosion, compaction, and other problems (Walker et al. 2009). Recovery of stable plant communities can be achieved by first stabilising the substrates with bioengineering techniques such as live stakes, brush layers, and wattle fences (Polster 2003; Singh 2010; Punetha et al. 2019), supplemented with establishment of early successional and mid-successional native plant cover and amendment of the soil with nutrients and organic matter. Successful growth of established plants on landslides may require different environmental conditions than colonising plants. The emphasis should be on assuring a diversity of functionally redundant species, so that if some plants do not survive, others with a similar life history can fill in the gaps (Walker et al. 2009). These species can stabilise and fertilise the surface and should have rapid growth and reproduction traits. On Peace River region landslides, native species to use in restoration plantings could include grasses such as blue wildrye (Elymus glaucus) and Canada bluejoint (Calamagrostis canadensis), forbs such as wild sarsaparilla (Aralia nudicaulis), palmate coltsfoot (Petasites frigidus), showy aster (Aster conspicuus) and false Solomon’s seal (Smilacina racemosa) and shrubs such as prickly rose (Rosa acicularis), saskatoon (Amelanchier alnifolia), alder species (Alnus spp.), red-osier dogwood (Cornus stolonifera), and highbush cranberry (Viburnum edule). Because the deposition zone is generally more stable, restoration efforts in this part of the slide should focus on management of species interactions, such as reducing competition between native species and exotics such as Melilotus officinalis. Management of interactions should retain some exotics for nitrogen fixing and nutrient input, but exotic densities should be reduced around native species. Restoration in the headscarp and chute zones should prioritise proactive erosion control and enhanced seed dispersal, perhaps using hydroseeding. In 109 addition, planting of native shrubs and tree seedlings could be done throughout the landslide wherever slopes are already stable, to diversify the species composition and increase the rate of succession. Two strategies to accelerate succession on the large bare or sparse zones and the grassdominated areas on the Beatton River and Cecil Lake landslides could be to increase natural dispersal by birds and amend the substrate surface with nutrients (Shiels and Walker 2001, 2003). The dispersal strategy with birds involves establishing perches in the open areas to encourage birds to fly closer to the centre of the landslide, where they may drop, regurgitate, or defecate seeds. The goal is to facilitate the establishment of native woody plants to shade out the graminoids and aggressive forbs. Success would depend on the bird species and the proximity of the perches from the forest edge. For seed deposition from birds to occur, there must be some vegetation on the slide (Shiels and Walker 2001). However, this poses a problem for restoration, because forest seed germination may be inhibited by competition with grasses, exotics, and other fast-growing pioneers. Forest seed germination may also be hindered on bare sites due to low organic matter and extreme microclimates. Commercial fertiliser and amendments with mature forest soil may help speed up plant colonisation. However, research has shown that organic matter input on landslides does not influence plant growth unless it is forest soil (Shiels et al. 2006). The type of organic substrate, frequency of organic deposition, and presence or absence of biota all directly influence soil nutrient patch dynamics, which ultimately affects plant development. Overall, restoration on landslides in the Peace River Region should focus on ecosystem recovery and biodiversity, rather than plant species composition. It is important to consider long-term recovery trends and effects of nonnative species succession because restoration 110 practices often incorporate nonnative species (Dale and Adams 2003). Further, the influence of invasive species and other disturbances needs to be fully integrated into successional theory. Biogeomorphic systems such as landslides are viewed as open and path-dependent systems by many researchers (Stallins 2006). Plant communities change as the substrate changes, and the communities also actively change the substrate. The material impacts of organisms serving as ecosystem engineers may be just as important as the trophic links, since they stabilise substrates, enhance weathering, provide habitat, and promote facilitation. There are many developmentally connected feedbacks between geomorphic and ecological components, and multiple causes and recurrences should be considered when studying succession and planning restoration projects. Further research on landslides in northern BC could enhance understanding of plant community development, geomorphic evolution, relationships between geomorphic diversity and vegetation diversity on landslides, and broader ecological processes on these disturbances. Resampling of vegetation plots on each of the three study areas after some years have passed could provide insights on whether species composition and site conditions are changing or becoming more diverse over time. Sampling on remote landslides away from the influence of cultivated fields or roadways would create a dataset for comparison of vegetation dynamics with solely native vegetation. In addition, a variety of landslides of different movement types, sizes, and ages could be sampled to provide a broader understanding of geomorphic, spatial, and temporal influences on alpha diversity. Experiments with bird perches, organic matter amendments, bioengineering, and hydroseeding on the bare slopes of the slides would present valuable lessons on the 111 feasibility and effectiveness of management options. Finally, detailed studies on wildlife habitat, wildlife use of landslide features, and connectivity to the surrounding undisturbed terrain would provide a perspective of the ecological role of landslides at the landscape level. All these findings could be applied more generally to our understanding of landslide ecology (Walker and Shiels 2013), and the options for management, and restoration here and elsewhere in the world. 112 2.7 References Avery, T.E. 1969. Forester’s guide to aerial photo interpretation. U.S. Department of Agriculture, Forest Service. Agriculture Handbook No. 308. Washington, D.C. 49 pp. Bazzaz, F. A. 1975. Plant species diversity in old‐field successional ecosystems in southern Illinois. Ecology, 56(2), 485-488. BC Gov (British Columbia Ministry of Forests and Range and British Columbia Ministry of Environment). 2010. Field manual for describing terrestrial ecosystems, 2nd edn. BCMFR Research Branch and BCMOE Resource Inventory Branch, Victoria, B.C. (Reprint with updates 2015.) BC Forest Service Research Branch. 2016. British Columbia plant species codes. BC Forest Service Research Branch. 2020. British Columbia plant species codes Version 14. Beaudry, L., R. Coupe, C. DeLong, and J. Pojar. 1999. Plant indicator guide for northern British Columbia: Boreal, Sub-Boreal, and Subalpine biogeoclimatic zones (BWBS, SBS, SBPS, and northern ESSF). Land Management Handbook 46, BC Ministry of Forests, Research Branch, Victoria, BC BECWeb. 2023. Biogeoclimatic Ecosystem Classification Program. Information website. https://www.for.gov.bc.ca/hre/becweb/resources/information-requests/index.html Belessiotis, V, and E. Delyannis. 2011. Solar drying. Solar Energy, 85: 1665-1691. Burton, P. 1991. Ecosystem restoration versus reclamation: the value of managing for biodiversity. Proceedings of the 15th Annual British Columbia Mine Reclamation Symposium in Kamloops, BC, 1991. The Technical and Research Committee on Reclamation. pp. 17-26. Burton, P.J., A.C. Balisky, L.P. Coward, S.G. Cumming, and D.D. Kneeshaw. 1992. The value of managing for biodiversity. The Forestry Chronicle 68(2): 225-237. DOI: 10.5558/tfc68225-2 Chao, A. 1984. Nonparametric estimation of the number of classes in a population. Scandinavian Journal of Statistics,11(4): 265-270. Chilton, R.R.H. 1981. A summary of climatic regimes of British Columbia. Ministry of Environment, Air Studies Branch, Assessment and Planning Division. 44 pp. 113 Chiu, C-H., and A. Chao. 2014. Distance-based functional diversity measures and their decomposition: a framework based on Hill numbers. PLoS ONE 9(7): e100014 1-17. Colwell, R.K., and J.A. Coddington. 1994. Estimating terrestrial biodiversity through extrapolation. Philosophical Transactions of the Royal Society of London B Biological Sciences, 345: 101-118. Cruden, D., and D. Varnes. 1996. Landslide types and processes. In: Turner, A., Schuster, R. (eds.), Special Report 247: Landslides Investigation and Mitigation. National Research Council, Transportation Research Board, Washington, DC, pp. 36-75. Dale, V., and W. Adams. 2003. Plant reestablishment 15 years after the debris avalanche at Mount St. Helens, Washington. The Science of the Total Environment, 313: 101-113. Dale, V. H., F.J. Swanson, and C.M. Crisafulli. 2005. Disturbance, survival, and succession: understanding ecological responses to the 1980 eruption of Mount St. Helens. In: Dale, V.H., F.J. Swanson, and C.M. Crisafulli (eds). Ecological Responses to the 1980 Eruption of Mount St Helens. Springer, New York, NY. https://doi.org/10.1007/0-387-28150-9_1. pp. 311 DeLong, C., A. Banner, W. H. Mackenzie, B. J. Rogers, and B. Kaytor. 2011. A field guide to ecosystem identification for the Boreal White and Black Spruce Zone of British Columbia. B.C. Ministry of Forests and Range, Forest Sciences Program, Victoria, B.C. Land Management Handbook No. 65. Demarchi, D.A. 2011. An introduction to the ecoregions of British Columbia. Third Edition. Ecosystem Information Section, Ministry of Environment, Victoria, B. C. 163 pp. Douglas, G.W., D.V. Meidinger, and J. Pojar (eds). 1998-2002. Illustrated Flora of British Columbia, volumes 1 to 8. B.C. Ministry of Environment, Lands and Parks, and B.C. Ministry of Forests, Victoria, B.C. FEIS. 2023. Fire Effects Information System. Syntheses about fire ecology and fire regimes in the United States. United States Forest Service, Department of Agriculture. Online source. https://feis-crs.org/feis/ Fisher, R.A., A.S. Corbet, and C.B. Williams. 1943. The relationship between the number of species and the number of individuals in a random sample of an animal population. Journal of Animal Ecology,12:42–58. 114 Foster, D. R., D.H. Knight, and J.F. Franklin. 1998. Landscape patterns and legacies resulting from large, infrequent forest disturbances. Ecosystems, 1: 497-510. Francescato, V., M. Scotton, D. Zarin, J. Innes, and D. Bryant. 2001. Fifty years of natural revegetation on a landslide in Franconia Notch, New Hampshire, U.S.A. Canadian Journal of Botany, 79: 1477-1485. Franklin, J.F., D. Lindenmayer, J.A. MacMahon, A. McKee, J. Magnuson, D.A. Perry, R. Waide, and D. Foster. 2000. Threads of continuity. Conservation Biology in Practice, 1(1): 816. Franklin, J. F., and J.A. MacMahon. 2000. Messages from a mountain. Science, 288(5469): 1183-1184. Geertsema, M., J.J. Clague, J.W. Schwab, and S.G. Evans. 2006. An overview of recent large catastrophic landslides in northern British Columbia, Canada. Engineering Geology, 83: 120-143. Geertsema, M., L. Highland, and L. Vaugeouis. 2009. Environmental impact of landslides. In Landslides – Disaster Risk Reduction. K. Sassa and P. Canuti (eds), Springer-Verlag. Berlin, Heidelberg. pp. 589-607. Geertsema, M., and J. Pojar. 2007. Influence of landslides on biophysical diversity – A perspective from British Columbia. Geomorphology, 89: 55-69. Global Mapper. 2020. Global Mapper v23 64 bit. Blue Marble Geographics. Goepel, K.D. 2012. Diversity Excel Template – Diversity Indices and True Diversity. Business Performance Management Singapore. https://bpmsg.com/diversity-18-12-12/ Good, I.J. 1953. The population frequencies of species and the estimation of population parameters. Biometrika, 40: 237-264. Gotelli, N., and R.K. Colwell. 2001. Quantifying biodiversity: procedures and pitfalls in the measurement and comparison of species richness. Ecology Letters, 4: 379-391. Gower, J.C. 1971. Statistical methods of comparing different multivariate analyses of the same data. In Hodson, F.R., D.G. Kendal, and P. Tautu, (eds) Mathematics in the archaeological and historical sciences. Edinburgh University Press, Edinburgh. pp. 138-149. 115 Haeussler, S., D. Coates, and J. Mather. 1990. Autecology of common plants in British Columbia: a literature review. Forest Resource Development Agreement (FRDA) report 158, 275 pp. Joint publication of BC Ministry of Forests, Research Branch and Forestry Canada, Pacific Forestry Centre, Victoria, B.C. Hill, O. 1973. Diversity and evenness: a unifying notation and its consequence. Ecology, 54(2):427-432. Holland, S.S. 1976. Landforms of British Columbia: A physiographic outline. British Columbia Geological Survey, British Columbia Department of Mines and Petroleum Resources, Mineralogical Branch, Province of British Columbia, Bulletin 48. 137 pp. Howes, D., and E. Kenk. 1997. Terrain classification system for British Columbia (version 2). BC Ministry of Environment, Recreational Fisheries Branch, and BC Ministry of Crown Lands, Surveys and Resource Mapping Branch, Victoria. Hurlburt, S.H. 1971. The non-concept of species diversity: a critique and alternative parameters. Ecology, 52: 577-586. Johnson, D., L. Kershaw, A. MacKinnon, J. Pojar, T. Goward, and D. Vitt. 1995. Plants of the western boreal forest and aspen parkland. Lone Pine Publishing and Canadian Forest Service, Edmonton, Alberta. 392 pp. Kershaw, L., A. MacKinnon, and J. Pojar. 1998. Plants of the Rocky Mountains. Lone Pine Publishing, Edmonton, Alberta. 384 pp. Klinkenberg, B. (editor). 2021. E-Flora BC: Electronic atlas of the plants of British Columbia. Department of Geography, University of British Columbia, Vancouver, BC. First Edition 2007. Kruskal, J.B. 1964. Multidimensional scaling by optimizing goodness of fit to a nonmetric hypothesis. Psychometrika 29(1): 1-27. Landau, F. 2004. Plant life forms (growth forms). Article retrieved online from landau.faculty.unlv.edu/lifeforms.htm Legendre, P., and L. Legendre. 1998. Numerical Ecology. Second English Edition. Elsevier, Amsterdam. 853 pp. Lloyd, M., and R.J. Ghelardi. 1964. A table for calculating the “equitability” component of species diversity. Journal of Animal Ecology, 33: 217-255. 116 Lord, T.M, and A.J. Green. 1986. Soils of the Fort St. John – Dawson Creek area, British Columbia. British Columbia Soil Survey, No. 42. Ottawa: Agriculture Canada Research Branch. 140 pp. MacArthur, R. H. 1965. Patterns of species diversity. Biological Reviews, 40: 510-533. MacArthur, R.H. 1972. Geographical Ecology. New York: Harper & Row. 269 pp. MacKenzie, W. 2012. Biogeoclimatic ecosystem classification of non-forested ecosystems in British Columbia. Province of British Columbia, Victoria, B.C. Technical Report 068 MacKenzie, W., and R. Klassen. 2004. VPro User Guide. British Columbia Ministry of Forests, Forest Science Program. Victoria, B.C. 106 pp. MacKenzie, W., and J.R. Moran. 2004. Wetlands of British Columbia: a guide to identification. Land Management Handbook No. 52. Research Branch, B.C. Ministry of Forests, Victoria, B.C. MacKinnon, A., J. Pojar, and R. Coupe. 1992. Plants of northern British Columbia. BC Ministry of Forests and Canadian Forest Resource Development Agreement. 352 pp. Magurran, A. 2004. Measuring biological diversity. Blackwell Publishing. Oxford, UK. 256 pp. Mather, P.M. 1976. Computational methods of multivariate analysis in physical geography. J. Wiley and Sons, London. 544 pp. Mathews, W.H. 1978. Quaternary stratigraphy and geomorphology of Charlie Lake (94A) map-area, British Columbia. Geological Survey of Canada, Ottawa. Paper 76-20. Mathews, W.H. 1980. Retreat of the last ice sheets in northeastern British Columbia and adjacent Alberta. Energy, Mines, and Resources Canada, Geological Survey of Canada, Bulletin 31, Ottawa. 28 pp. McCune, B., J.B. Grace, and D.L. Urban. 2002. Analysis of Ecological Communities. MjM Software Design, Gleneden Beach, OR. 300 pp. McCune, B., and D. Keon. 2002. Equations for potential annual direct incident radiation and heat load. Journal of Vegetation Science, 13(4): 603-606. McCune, B., and M.J. Mefford. 2016. PC-ORD. Multivariate analysis of Ecological Data, Version 7.0 for Windows. Wild Blueberry Media, Corvallis, Oregon, U.S.A. McIntosh, R.P. 1967. An index of diversity and the relation of certain concepts to diversity. Ecology, 48: 392-404. 117 Meidinger, D., and J. Pojar (eds). 1991. Ecosystems of British Columbia. Special Report Series No. 6. BC Ministry of Forests, Victoria, BC. Microsoft Corporation. 2010. Microsoft 365: Microsoft Excel. Retrieved from: https://office.microsoft.com/excel Morris, E.K., T. Caruso, F. Buscot, M. Fischer, C. Hancock et al. 2014. Choosing and using diversity indices: insights for ecological applications from the German Biodiversity Exploratories. Ecology and Evolution, 4(18): 3514-3524. Motomura, I. 1932. On the statistical treatment of communities. Zoological Magazine Tokyo, 44: 379-383. Nagendra, H. 2002. Opposite trends in response for the Shannon and Simpson indices of landscape diversity. Applied Geography, 22: 175-186. Noss, R.F. 1990. Indicators for monitoring biodiversity: a hierarchical approach. Conservation Biology, 4(4): 355-364. Oertli, B., D.A. Joye, E. Castella, R. Juge, D. Cambin, and J.-B. Lachavanne. 2002. Does size matter? The relationship between pond area and biodiversity. Biological Conservation, 104: 59-70. Ortiz-Burgos, S. 2016. Shannon-Weaver Diversity Index. In: Kennish, M.J. (eds) Encyclopedia of Estuaries. Encyclopedia of Earth Sciences Series. Springer, Dordrecht. https://doi.org/10.1007/978-94-017-8801-4_233 PC-ORD. 2015. PC-ORD website. https://www.pcord.com/ Peck, J.E. 2016. Multivariate Analysis for Ecologists: Step-by-Step, Second edition. MjM Software Design, Gleneden Beach, OR. 192 pp. Peet, R.K. 1974. The measurement of species diversity. Annual Review of Ecology and Systematics, 5: 285-307. Phillips, J., and C. Lorz. 2008. Origins and implications of soil layering. Earth-Science Reviews, 89: 144-155. Pickett, S., M. Cadenasso, and S. Meiners. 2009. Ever since Clements: from succession to vegetation dynamics and understanding to intervention. Applied Vegetation Science, 12(1): 9-21. 118 Pielou, E.C. 1966. The measurement of diversity in different types of biological collections. Journal of Theoretical Biology, 13: 131-144. Pitkanen, S. 2000. Classification of vegetational diversity in managed boreal forests in eastern Finland. Plant Ecology, 146(1): 11-28. Pojar, J., K. Klinka, and D.V. Meidinger. 1987. Biogeoclimatic ecosystem classification in British Columbia. Forest Ecology Management, 22: 119-154. Polster, D. F. 2003. Soil bioengineering for slope stabilization and site restoration. Paper presented Sudbury Mining and the Environment III, May 25-28. Available online at https://pdf.library.laurentian.ca/medb/conf/Sudbury03/Amendments/122.pdf Punetha, P., M. Samanta, and S. Sarkar. 2019. Bioengineering as an effective and ecofriendly soil slope stabilization method: A review. In: Prahdan,S., V. Vishal, T. Singh (eds) Landslides: Theory, practice and modelling. Advances in Natural and Technological Hazards Research, vol 50. Springer, Cham. pp. 201-224. Restrepo, C., L. Walker, A. Shiels, R. Bussmann, L. Claessens, S. Fisch, P. Lozano, G. Negi, L. Paolini, G. Poveda, C. Ramos-Scharrón, M. Richter, and E. Velázquez. 2009. Landsliding and its multiscale influence on mountainscapes. BioScience, 59(8): 685-698. Ricotta, C., and G. Avena. 2003. On the relationship between Pielou’s evenness and landscape dominance within the context of Hill’s diversity profiles. Ecological Indicators, 2: 361-365. Rozycka, M., P. Migon, and A. Michniewicz. 2016. Topographic Wetness Index and Terrain Ruggedness Index in geomorphic characterisation of landslide terrains, on examples from the Sudetes, SW Poland. Zeitschrift fur Geomorphologie, Supplementary Issue, 1-20. Saito, H., S. Uchiyama, and K. Teshirogi. 2021. Rapid vegetation recovery at landslide scars detected by multitemporal high-resolution satellite imagery at Aso volcano, Japan. Geomorphology, https://doi.org/10.1016/j.geomorph.2021.107989 Sayn-Wittgenstein, L. 1960. Recognition of tree species on air photographs by crown characteristics. Forest Research Division Technical Note 95. Department of Forestry, Forest Research Laboratory, Government of Canada, Ottawa, Ontario. 56 pp. Sayn-Wittgenstein, L. 1978. Recognition of tree species on aerial photographs. Information Report FMR-X-118. Forest Management Institute, Canadian Forest Service, Department of the Environment, Ottawa. 106 pp. 119 Shannon, C.E. 1948. A mathematical theory of communication. Bell System Technical Journal. 27: 379-423. Doi: 10.1002/j.1538-7305.1948.tb01338.x. Shiels, A. 2006. Leaf litter decomposition and substrate chemistry of early successional species on landslides in Puerto Rico. Biotropica, 38(3): 348-353. Shiels, A., and L. Walker. 2001. Accelerating plant colonisation on landslides in Puerto Rico by additions of bird perches and organic matter. Tropical Ecosystems: Structure, Diversity and Human Welfare. Proceedings of the International Conference on Tropical Ecosystems. K.N. Ganeshaiah, R. Uma Shaanker and K.S. Bawa (eds). Published by Oxford-IBH, New Delhi. pp. 661-664. Shiels, A., and L. Walker. 2003. Bird perches increase forest seeds on Puerto Rican landslides. Restoration Ecology, 11(4): 457-465. Shiels, A., L. Walker, and D. Thompson. 2006. Organic matter inputs create variable resource patches on Puerto Rican landslides. Plant Ecology, 184: 223-236. Simpson, E.H. 1949. The measurement of diversity. Nature 163:688. Singh, A. K. 2010. Bioengineering techniques of slope stabilization and landslide mitigation. Disaster Prevention and Management: An International Journal, 19(3): 384-397. DOI:10.1108/09653561011052547 Smith, R. 1986. Soils, vegetation, and forest growth on landslides and surrounding logged and old-growth areas on the Queen Charlotte Islands. Land Management Report 41. BC Ministry of Forests, Victoria, BC. Soil Classification Working Group. 1998. The Canadian system of soil classification. Third edition. Agriculture and Agri-Food Canada, Research Branch, Publication 1646, 187 pp. Sokal, R.R., and F.J. Rohlf. 1995. Biometry: The Principles and Practice of Statistics in Biological Research. Third Edition, W.H. Freeman and Col., New York. 887 pp. Sousa, W. P. 1984. The role of disturbance in natural communities. Annual Review of Ecology and Systematics, 15(1), 353-391. Stallins, J.A. 2006. Geomorphology and ecology: Unifying themes for complex systems in biogeomorphology. Geomorphology, 77(3-4):207-216. 120 Takaoka, S. 2015. Origin and geographical characteristics of ponds in a high mountain region of central Japan. Limnology, 16:103-112. Tokeshi, M. 1993. Species abundance patterns and community structure. Advances in Ecological Research, 24:112-186. Turner, M., W. Baker, C. Peterson, and R. Peet. 1998. Factors influencing succession: lessons from large, infrequent natural disturbances. Ecosystems, 1(6): 511-523. Valentine, K.W.G. 1978. The soil landscapes of British Columbia. Ministry of Environment, Resource Analysis Branch, Victoria, BC. 197 pp. Walker, L., and R. del Moral. 2008. Lessons from primary succession for restoration of severely damaged habitats. Applied Vegetation Science, 12(1): 55-67. Walker, L., and A. Shiels. 2013. Landslide Ecology. Cambridge University Press. New York, NY. 300 pp. Walker, L., E. Velazquez, and A. Shiels. 2009. Applying lessons from ecological succession to the restoration of landslides. Plant and Soil, 324(1-2): 157-168. Wallis, C.I.B., Y.C. Tiede, E. Beck, K. Bohning-Gaese, R. Brandl et al. 2021. Biodiversity and ecosystem functions depend on environmental conditions and resources rather than the geodiversity of a tropical biodiversity hotspot. Scientific Reports 11: 24530. Whittaker, R. 1960. Vegetation of the Siskiyou Mountains, Oregon and California. Ecological Monographs, 30:279-338. Whittaker, R. H. 1965. Dominance and diversity in land plant communities. Science, 147: 250-260. Zaiontz, C. 2017. “Two Sample Kolmogorov-Smirnov Table.” Real Statistics Using Excel. Online source: real-statistics.com/non-parametric-tests/goodness-of-fit-tests/two-samplekolmogorov-smirnov-test/ 121 Chapter 3. Beta diversity as a measure of biophysical turnover on landslides 3.1 Introduction and Background Through marked alteration of vegetation and environment, landslides create diversity on a landscape. Diversity in nature can be described in a series of scales forming a hierarchy. As outlined in Chapter 2, the three basic levels of diversity are alpha diversity, beta diversity, and gamma diversity (Whittaker 1960). These three scales of diversity are related, and often quantification of one level is required before quantifying another. Alpha and gamma diversity refer to defined geographic units and are classified as inventory diversity (Whittaker 1972). Alpha diversity is within-site diversity, while gamma diversity is described at a regional level. These two diversities may be measured in similar ways, although they are not transferable (Tuomisto 2010a). Whittaker coined the concept of a third category of diversity, namely beta diversity. Beta diversity is generally understood as diversity between sites or communities. It refers to variation between samples and is classified as differentiation diversity. It requires a different set of techniques for measurement than inventory diversity. Essentially, beta diversity is assessed in relation to complementarity or relative similarity or dissimilarity between communities (Magurran 2004). Beta diversity represents a key aspect of the spatial pattern of biodiversity. Whittaker (1960) described beta diversity in its simplest form as the change in species composition over an area, or the extent of change of community composition in relation to a gradient or pattern of environments. For this reason, it has also been referred to as spatial turnover. Generally, beta diversity increases as similarity in species composition decreases. A landscape with a certain number of species and little overlap in species composition over an area will be more diverse than an equivalent landscape with equally speciose assemblages but many shared 122 species from site to site. Beta diversity has been found to be more effective than alpha diversity at detecting change in ecological communities (Seguin et al. 2013), and therefore is valuable in assessing the effects of disturbances over time as well as space. 3.1.1 Measurement of beta diversity Unfortunately, many different definitions and concepts of beta diversity have arisen over the years, accompanied by an equally varying number of methods to describe and quantify it. This lack of standardisation has been the source of debate and discussion in recent years (Koleff et al. 2003; Moreno and Rodriguez 2010; Tuomisto 2010a; Tuomisto 2010b; Tuomisto 2010c; Anderson et al. 2011). Koleff et al. (2003) noted at least 24 measures of beta diversity for presence/absence data. All the variants of beta diversity measure some sort of differentiation or heterogeneity, but they each represent very different phenomena (Tuomisto 2010a). The many different measurements of these variants cannot be compared numerically. The debate over a lack of a unified concept and measure has led some to call beta diversity a key concept in ecology, while others deem it of no use (Jurasinski et al. 2009). In this current study, beta diversity refers to the change in species composition over space, or species turnover, and will focus on turnover as it relates to distance or area. There are two main categories of beta diversity measures: species richness (coefficients that examine variation in richness across scales) and species composition (coefficients that examine variation in species composition between samples) (Jurasinski et al. 2009). Species richness measures assess the extent of difference between two or more areas of alpha diversity relative to gamma diversity, where gamma diversity is usually measured as total species richness. Species composition measures focus on the differences in species 123 composition amongst areas of alpha diversity and were designed to measure complementarity or similarity/dissimilarity (Barwell et al. 2015; Chao and Chiu 2016). They employ similarity/dissimilarity coefficients, slope of distance-decay relationships, sum of squares species matrices, or gradient length in ordination space techniques. Species composition measures essentially assess the biotic distinctness of assemblages and include the Jaccard and Bray-Curtis coefficients. A third category of measures use the species-area relationship to measure turnover related to species accumulation with area. Beta diversity indices As with alpha diversity, various indices have been devised for beta diversity. Most indices use presence/absence data and so focus on the species richness component of diversity. One of the simplest and most effective measures is Whittaker’s (1960) measure Bw, which is obtained by dividing total species recorded in the system by average sample diversity (species richness). Of six beta diversity measures tested by Wilson and Shmida (1984) for effectiveness in measuring community turnover, Whittaker’s measure was found to fulfill most of the authors’ criteria of number of community (assemblage) changes, additivity, independence from alpha diversity, and independence from excessive sampling, with fewest restrictions. The more complementary two sites are, the higher their beta diversity. Complementarity describes the difference between sites in terms of the species they support. The beta diversity of pairs of sites can be described by using a similarity/dissimilarity coefficient. A matrix can be constructed using the Jaccard (1908) similarity index, which tallies species gained and species lost. 124 The Sorensen (1948) measure is another similarity measure that is popular and viewed as very effective. It uses presence/absence and is identical to the Bray-Curtis presence/absence coefficient. One disadvantage of this measure is that if samples differ markedly in species richness, the Sorensen measure will always be large (Lennon et al 2001). However, it can be modified to accommodate quantitative data (Bray and Curtis 1957). A significant advantage of the beta diversity measures discussed is their ease of calculation and application. However, the coefficients do not take into consideration relative abundance of species. Dominant species have no more weight in a presence/absence beta diversity measure than a species represented by one individual. This has resulted in the development of similarity/dissimilarity measures based on quantitative data, such as the Bray-Curtis index (Bray and Curtis 1957). 3.1.2 Landslide beta diversity Landslides create harsh environments that can act as ecological filters for plant community development (Furusawa et al. 2023). Because they are large-scale disturbances caused by stochastic events, landslides are difficult to study and generalise. The environment that results from a landslide often consists of a patchwork of rafted materials amid a variety of exposed and deposited substrates, which can lead to increased patchiness across the landscape. This greater patchiness presumably translates to increased beta diversity. Community assembly occurs by both stochastic/random (i.e. ecological drift) and deterministic (i.e. niche selection) processes. General patterns of community responses to landslides and environmental influences on community assemblages after a landslide are poorly understood. The extreme environment following a landslide is expected to favour 125 niche-assembled communities comprised of specialist species, but little is known about this process. Disturbances such as landslides can alter beta diversity by changing the relative importance of community assembly mechanisms that influence clustering of species across landscapes (Myers et al. 2015). Disturbance can increase clumping either through divergent selection of niches across environmental gradients or through reduced dispersal if species become rare after disturbance, or by a combination of the two processes. Disturbed landscapes may thus have a lower species richness and a lower overall alpha diversity than undisturbed landscapes, yet greater beta diversity. In this chapter, I attempt to answer the question: To what extent does turnover or beta diversity in microsites and plant species occur on landslides? I also investigate whether vegetation beta diversity is correlated in any way with site beta diversity. For both lines of investigation, I also assess whether there is a difference between different sizes or ages of landslides and between landslide and undisturbed terrain study sites. 3.1.3 Study areas The sampling transects for this study were established on the same three study areas described in Chapter 2, in the Peace River Region of northeastern BC on landslides < years old in glaciolacustrine material. The three landslides were Beatton River, Cecil Lake, and Hasler Flats. The Beatton River landslide most recently activated in 2015 and was approximately 30 ha. The Cecil Lake landslide occurred in 1998 and was about 56 ha. The much smaller Hasler Flats landslide occurred in 2013 and was only 1.5 ha in size. All three landslides initiated on plateaus and travelled downslope into rivers or streams. Detailed 126 information on each landslide and its associated surrounding terrain can be found in Chapter 2 on alpha diversity, in Section 2.2. 3.2 Methods 3.2.1 Data collection Sampling of vegetation cover and environment variables was carried out on paired landslide and undisturbed study sites in the Peace River Region of British Columbia on the three study areas: Beatton River, Cecil Lake, and Hasler Flats. On the Beatton River and Cecil Lake landslide study sites, seven randomly located transects were sampled, comprised of up to 30 m of a series of 1 x 1 m quadrats (plots). On the Hasler Flats landslide study site, just three transects were established, due to the much smaller size of the landslide (1.5 ha). In total, 17 landslide transects were sampled. On the associated undisturbed study sites surrounding each landslide, three transects were sampled for each of the three study areas, for a total of nine undisturbed transects. Transect commencement points for each transect were randomly located on a grid in the office prior to beginning field work, with point locations and transect directions determined using an online random number generator. To further ensure randomness of data, field sampling was done to the right of the transect defined by a 30 m tape measure on even numbered transects and to the left of the transect on odd numbered transects. Individual quadrats were delineated using a 1 x 1 m square plot made of PVC (polyvinyl chloride) tubing (Figure 3-1). At each plot, plant species and associated percent covers were recorded, along with ground substrate percentages as per Land Management Handbook 25 (BC Gov 2010), as well as predominant slope and predominant aspect. Slope and aspect were measured at the microsite level (i.e. the 1 x 1 m 127 quadrat plot). Aspect was eventually converted to folded aspect values along the north-south access to facilitate its analysis as a continuous variable (McCune and Keon 2002). Any vegetation or site type changes along the transect were recorded. Adjacent plots were added successively along the transect until two plots were sampled consecutively with no new species or the 30 m transect ended, whichever came first. If plots fell on a transition between two visibly different vegetation or substrate types, the plot was moved to the next metre marker where it was completely within the new type. Additionally, if there were no new species between two consecutive plots but there was a type change further along the 30 m of the transect, the next plot was established at the next metre marker fully within the new type. The number of quadrats per landslide transect ranged from 15-30 for Beatton River and 1430 for Cecil Lake, while all transects for Hasler Flats had 30 quadrats. All undisturbed transects for Beatton River and Hasler Flats had 30 quadrats, while for Cecil Lake the number of quadrats per undisturbed transect ranged from 28-30. Sampling intensity for the landslide study sites was 0.23 transects/ha and 5.63 quadrats/ha for Beatton River, 0.12 transects/ha and 3.00 quadrats/ha for Cecil Lake, and 2.00 transects/ha and 60 quadrats/ha for Hasler Flats. Sampling intensity for the undisturbed study sites worked out to 0.1 transects/ha and 3.00 quadrats/ha for Beatton River, 0.05 transects/ha and 1.57 quadrats/ha for Cecil Lake, and 2.00 transects/ha and 60 quadrats/ha for Hasler Flats. For all transects, representative photographs were taken at various locations along the length. 128 Figure 3-1. Sampling transects with a series of 1 x 1 m quadrats along a measuring tape. Image on the left is Beatton River undisturbed transect BEtu1a. (Photo August 26, 2018) Image on the right is Beatton River landslide transect BEtl1a. (Photo July 18, 2017, by P. Burton. Used with permission.) 3.2.2 Data analysis Vegetation data for each transect was entered into Excel spreadsheets, formatted, and then exported to the multivariate analysis program PC-ORD v. 7.10 (McCune and Mefford 2016; McCune and Mefford 2018) for analysis. For plots with no vegetation or 100 percent water, separate columns were created titled “NO_PLANTS” and “WATER”. Two transects on the Beatton River landslide had no vegetation the entire length of the transect, and thus were eliminated from the PCORD analysis because it was discovered after field sampling that a beta diversity value could not be calculated in PCORD when empty rows were present. For 129 each remaining transect on each landslide and in the associated surrounding undisturbed terrain, an initial value of Whittaker’s measure of beta diversity (Bw) was calculated in PCORD, using the formula Bw = Gamma (overall) diversity/Alpha (local) diversity. Environment data for each transect was entered into Excel spreadsheets and formatted for analysis in PC-ORD. Slope and folded aspect were inputted in separate columns. Folded aspect was calculated in Excel using the following formula: ABS(180 - ABS(aspect -225°) where ABS refers to the absolute value, and aspect was measured in degrees azimuth from true north. Folding aspect around the southwest slope-facing direction is considered representative of heat intensity on a site in the northern hemisphere (McCune and Keon 2002). Substrate values for organic matter, decaying wood, bedrock, rock (cobbles and stones), mineral soil, and water were inputted in separate columns. Once the environment data was formatted, each transect dataset was imported into PC-ORD and Whittaker’s beta diversity Bw was calculated. Descriptive statistics were then calculated in Excel for the vegetation and environment beta diversity data for each study site. To provide background information and a fuller picture of diversity on the sites, alpha diversity values for vegetation and environment variables were calculated for each transect by averaging the values of all quadrats/plots along the transect. Richness, evenness, Shannon index (H’), and Simpson’s index (1-D) were all calculated for each transect in PC-ORD. The higher the evenness of a sample, the more similar the abundances are to each other in an area, and the higher the alpha diversity. The Shannon index incorporates richness and evenness to quantify the uncertainty in predicting the identity of an individual taken at random from a 130 dataset. It is related to the weighted geometric mean of the proportional abundances (evenness). The higher the Shannon index, the greater the uncertainty. Simpson’s index of diversity ranges between 0 and 1 and represents the probability that two individuals randomly selected from a sample will belong to different groupings. The higher the Simpson value, the greater the sample alpha diversity. Descriptive statistics were calculated in Excel for the vegetation and environment alpha diversity data for each study site, using the individual transect data generated in PC-ORD. A scatter graph was created to display the relationship between the resultant vegetation beta diversity values and environment beta diversity values for all landslide transects, and for all undisturbed transects. Regression lines of vegetation beta diversity as a function of environment beta diversity and associated regression statistics were then calculated, displayed, and compared for the landslide and undisturbed transects. 3.3 Results 3.3.1 Transect vegetation beta diversity 3.3.1.1 Landslide transect vegetation beta diversity The results for the landslide transect vegetation beta diversity calculations in PC-ORD revealed a range of values, both within and among study sites (Table 3-1). 131 Table 3-1. Landslide transect vegetation beta diversity (Bw) values. Transect Be ta dive rsity (Bw) Be atton River Mean Bw BEtl1a 2.7 BEtl1 1.8 BEtl4 0.8 2.42 BEtl5 3.0 BEtl6 3.8 Cecil Lake CEtl1 1.9 CEtl2 2.3 CEtl3c 5.5 CEtl4 2.8 3.10 CEtl5 2.3 CEtl6 4.2 CEtl7 2.7 Hasler Flats HAtl1a 2.9 HAtl2 3.0 3.17 HAtl3 3.6 Ove rall (all transects) 2.89 SD CV % 1.15 47.69 1.29 41.52 0.38 11.96 1.11 38.39 Cecil Lake transects showed the greatest range of landslide vegetation beta diversity (Bw) values, while Hasler Flats transects had the smallest range. Cecil Lake also yielded the single transect with the highest beta diversity value (CEtl3c at 5.5), whereas Beatton River had the transect with the lowest beta diversity value (BEtl4 at 0.8) of all the landslide transects. Hasler Flats presented the highest mean landslide transect vegetation Bw (3.17) whereas Beatton River had the lowest mean Bw (2.42) of the three study sites. Cecil Lake showed the highest standard deviation (SD), Beatton River had the highest coefficient of variation (CV), and Hasler Flats exhibited the lowest SD and CV. Both Hasler Flats and Cecil Lake had a higher mean than the overall mean Bw value for all landslide vegetation transects in the study. 132 For background information, alpha diversity values for vegetation were calculated for each landslide transect study site (Table 3-2). Mean richness, evenness, Shannon index (H’), and Simpson’s index (D) were all calculated in Excel from individual transect values generated in PC-ORD. Although these values were generally lower than those calculated for the landslide relevés in Chapter 2, the overall trend was the same for ranking the landslides. Table 3-2. Landslide transect vegetation alpha diversity statistics. Diversity Measure Beta diversity (Bw) Richness (S) Eve nness (E) Shannon (H') Simpson (D) Mean 2.42 6.64 0.58 0.96 0.46 Beatton River SD CV % 1.15 47.69 2.61 39.32 0.09 15.41 0.37 38.54 0.15 32.54 Mean 3.1 11.4 0.67 1.55 0.66 Cecil Lake SD CV % 1.29 41.52 2.99 26.26 0.05 6.73 0.39 25.05 0.15 22.55 Mean 3.17 12.3 0.63 1.47 0.64 Hasler Flats SD CV% 0.38 11.96 1.57 12.78 0.11 18.28 0.22 15.05 0.07 11.22 Overall - All landslide transects Mean SD CV % 2.89 1.11 38.39 9.99 3.51 35.08 0.63 0.08 13.06 1.34 0.43 32.41 0.59 0.16 27.07 Of the three landslides, Hasler Flats had the highest vegetation richness (12.3) while Beatton River had the lowest richness (6.64) but the highest CV (39.32%). Hasler Flats and Cecil Lake mean richness values were higher than the overall mean. Cecil Lake showed the highest evenness while Hasler Flats yielded the lowest evenness value. At the same time, Hasler Flats had the highest CV. The Shannon index was highest on the Cecil Lake landslide and lowest on the Beatton River landslide. Beatton River showed the highest CV for Shannon index. Cecil Lake and Hasler Flats both had a mean Shannon index that was higher than that of the mean overall value for all landslide transects. Cecil Lake also yielded the highest Simpson index while Beatton River was the lowest. Cecil Lake and Hasler Flats both had a higher mean Simpson index than the overall mean. 3.3.1.2 Undisturbed transect vegetation beta diversity Table 3-3 shows the vegetation beta diversity values calculated in PC-ORD for each undisturbed transect. 133 Table 3-3. Undisturbed transect vegetation beta diversity (Bw) values. Transect Be ta dive rsity (Bw) Be atton River Mean Bw BEtu1a 3.3 BEtu2b 1.8 2.50 BEtu3a 2.4 Ce cil Lake CEtu1 2.4 CEtu2 3.2 2.87 CEtu3a 3.0 Hasler Flats HAtu1a 2.3 HAtu2 1.4 2.03 HAtu3 2.4 Ove rall (all transects) 2.47 SD CV % 0.75 30.20 0.42 14.52 0.55 27.09 0.63 25.40 Of the three undisturbed study sites, Beatton River yielded the greatest range of transect vegetation Bw values while Cecil Lake showed the smallest range. Beatton River contained the undisturbed transect with the overall highest Bw value (BEtu1a at 3.3), while Hasler Flats had the transect with the lowest Bw value (HAtu2 at 1.4). For the undisturbed transect vegetation, Hasler Flats yielded the lowest mean Bw, at 2.03. Cecil Lake presented the highest mean Bw, at 2.87. Cecil Lake also had the lowest SD and CV. Both Beatton River and Cecil Lake showed a mean Bw that was higher than the overall mean for all the undisturbed transects. Beatton River also had a higher SD and CV than these overall values for all undisturbed transects. For background information, alpha diversity values for vegetation were calculated for each undisturbed transect study site (Table 3-4). Mean richness, evenness, Shannon index (H’), and Simpson’s index (D) were all calculated in Excel from the individual transect values generated in PC-ORD. Although these alpha diversity values were for the most part lower 134 than those calculated for the undisturbed relevés in Chapter 2, the results ranked the undisturbed study sites in the same order. Table 3-4. Undisturbed transect vegetation alpha diversity statistics. Diversity Me asure Beta diversity (Bw) Richness (S) Evenness (E) Shannon (H') Simpson (D) Mean 2.50 10.50 0.73 1.69 0.73 Beatton River SD CV % 0.75 30.20 1.15 10.98 0.05 7.43 0.19 11.44 0.06 8.36 Mean 2.87 10.17 0.70 1.56 0.69 Cecil Lake SD CV % 0.42 14.52 1.89 18.57 0.04 6.33 0.27 17.60 0.09 12.76 Mean 2.03 16.53 0.74 2.04 0.80 Hasler Flats Overall - All undisturbed transects SD CV % Mean SD CV % 0.55 27.09 2.47 0.63 25.40 1.50 9.06 12.40 3.38 27.25 0.04 5.10 0.72 0.04 6.10 0.15 7.14 1.77 0.28 16.05 0.04 4.94 0.74 0.08 10.25 Hasler Flats showed the highest mean undisturbed transect vegetation richness value (16.53), while Cecil Lake had the lowest richness value (10.17). Only Hasler Flats exhibited a mean richness higher than the overall mean richness for all undisturbed study sites. Regarding evenness, all three undisturbed study sites were similar in mean value, but Beatton River yielded the highest CV while Hasler Flats had the lowest CV. The Shannon index was highest at Hasler Flats and lowest at Cecil Lake, but Cecil Lake had the highest SD and CV. Only Hasler Flats showed a higher Shannon index than the overall mean. Hasler Flats also had the highest Simpson index, while Cecil Lake yielded the lowest Simpson index. Additionally, Cecil Lake had the highest SD and CV for Simpson’s index. 3.3.2 Transect environment beta diversity The results for the environment beta diversity calculations in PC-ORD generally showed a smaller range of values as well as lower beta diversity values overall, compared to the vegetation beta diversity. 3.3.2.1 Landslide transect environment beta diversity Table 3-5 presents the beta diversity values calculated in PC-ORD for landslide transect environment variables. 135 Table 3-5. Landslide transect environment beta diversity (Bw) values. Transect Be ta dive rsity (Bw) Be atton River Mean Bw BEtl1a 0.2 BEtl1 0.2 BEtl4 0.0 0.22 BEtl5 0.1 BEtl6 0.6 Cecil Lake CEtl1 0.2 CEtl2 0.2 CEtl3c 0.9 CEtl4 0.3 0.43 CEtl5 0.2 CEtl6 0.9 CEtl7 0.3 Hasler Flats HAtl1a 0.6 HAtl2 0.3 0.43 HAtl3 0.4 Ove rall (all transects) 0.36 SD CV % 0.23 103.65 0.33 75.87 0.15 35.25 0.27 75.56 The Cecil Lake study site showed the greatest range in landslide transect environment Bw values while Hasler Flats had the smallest range. Beatton River contained the transect with the overall lowest Bw (BEtl4, at 0.0), while Cecil Lake contained two transects with the highest Bw (CEtl3c and CEtl6, at 0.9). Hasler Flats yielded the highest mean landslide transect environment Bw, at 0.43, while Beatton River had the lowest mean Bw, at 0.22. Hasler Flats showed the lowest SD and CV. Both Hasler Flats and Cecil Lake had mean Bw values that were higher than the overall mean environment Bw value for all landslide transects. For background information, alpha diversity values for environment variables were calculated for each landslide transect study site (Table 3-6). Mean richness, evenness, 136 Shannon index (H’), and Simpson’s index (D) were all calculated in Excel using the individual transect values generated in PC-ORD. Alpha diversity values were generally lower than those calculated for landslide geotypes in Chapter 2, but the results overall ranked the landslide study sites in the same order. Table 3-6. Landslide transect environment alpha diversity statistics. Diversity Measure Beta diversity (Bw) Richness (S) Evenness (E) Shannon (H') Simpson (D) Mean 0.22 4.42 0.67 1.00 0.55 Beatton River SD CV % 0.23 103.65 0.30 6.86 0.05 6.79 0.11 11.28 0.05 8.60 Mean 0.43 3.87 0.69 0.92 0.52 Ce cil Lake SD CV % 0.33 75.87 0.17 4.40 0.08 11.03 0.12 13.23 0.07 14.05 Mean 0.43 4.20 0.63 0.90 0.48 Hasler Flats Overall -All landslide transects SD CV % Mean SD CV % 0.15 35.25 0.36 0.27 75.56 0.50 11.90 4.12 0.37 9.04 0.04 7.13 0.67 0.06 9.36 0.14 15.13 0.94 0.12 12.79 0.06 11.90 0.52 0.06 12.11 Beatton River showed the highest landslide transect environment richness while Cecil Lake had the lowest richness. Hasler Flats yielded the highest CV, while Cecil Lake had the lowest CV. Both Beatton River and Hasler Flats showed a higher mean environment richness than the overall average for all landslide transects. Of the three study sites, Cecil Lake had the highest environment evenness, while Hasler Flats exhibited the lowest evenness. Cecil Lake also showed the highest CV. For landslide environment Shannon diversity, Beatton River had the highest value, while Hasler Flats yielded the lowest Shannon index. However, Hasler Flats also had the highest CV. Only Beatton River showed a higher mean Shannon than the overall mean Shannon value. For Simpson diversity, Beatton River had the highest value while Hasler Flats showed the lowest value. Beatton River was higher than the mean overall Simpson value, while Cecil Lake was tied with the overall mean. 3.3.2.2 Undisturbed transect environment beta diversity Table 3-7 shows the environment beta diversity values for each undisturbed transect. 137 Table 3-7. Undisturbed transect environment beta diversity (Bw) values. Transect Be ta dive rsity (Bw) Be atton River Mean Bw BEtu1a 0.4 BEtu2b 0.4 0.30 BEtu3a 0.1 Ce cil Lake CEtu1 0.1 CEtu2 0.1 0.13 CEtu3a 0.2 Hasler Flats HAtu1a 0.2 HAtu2 0.4 0.37 HAtu3 0.5 Ove rall (all transects) 0.27 SD CV % 0.17 57.74 0.06 43.30 0.15 41.66 0.16 59.29 Beatton River and Hasler Flats were tied for the highest range of undisturbed transect environment Bw values, while Cecil Lake had the lowest range of Bw values. Hasler Flats yielded the highest mean Bw value of the three undisturbed transect environment data sets, whereas Cecil Lake showed the lowest Bw. Beatton River had the highest CV, while Hasler Flats had the lowest CV. Of the three study sites, only Hasler Flats exhibited a higher mean environment Bw than the overall mean value for all undisturbed transects. For background information, alpha diversity values for environment variables were calculated for each undisturbed transect study site (Table 3-8). Mean richness, evenness, Shannon index (H’), and Simpson’s index (D) were all calculated in Excel using diversity values that were generated in PC-ORD for each transect. These alpha diversity values were overall lower than those calculated for undisturbed geotypes in Chapter 2, and for some indices the undisturbed study sites were ranked differently than in Chapter 2. 138 Table 3-8. Undisturbed transect environment alpha diversity statistics. Diversity Measure Beta diversity (Bw) Richness (S) Eve nne ss (S) Shannon (H') Simpson (D) Mean 0.30 3.60 0.81 1.03 0.60 Beatton River SD CV % 0.17 57.74 0.00 0.00 0.07 8.64 0.09 8.74 0.04 6.24 Me an 0.13 3.53 0.65 0.81 0.49 Cecil Lake SD CV % 0.06 43.30 0.21 5.89 0.04 5.80 0.02 2.46 0.03 6.26 Mean 0.37 3.47 0.71 0.87 0.53 Hasler Flats Overall - All undisturbed transe cts SD CV % Mean SD CV % 0.15 41.66 0.27 0.16 59.29 0.21 6.00 3.53 0.16 4.47 0.05 6.98 0.72 0.08 11.64 0.10 11.66 0.90 0.12 13.14 0.04 7.31 0.54 0.06 10.47 For the undisturbed transect environment alpha diversity data, Beatton River showed the highest environment richness whereas Hasler Flats had the lowest richness. Hasler Flats yielded the highest richness CV, while Beatton River had the lowest CV. The Beatton River richness value was also higher than the mean overall richness, while richness for Cecil Lake was tied with the overall mean. Beatton River showed the highest evenness, while Cecil Lake had the lowest evenness. Only Beatton River had a higher evenness than the overall average for all undisturbed transects. Beatton River also yielded the highest Shannon and Simpson diversities, while Cecil Lake had the lowest values. Beatton River was the only study site with environment Shannon and Simpson diversities higher than the overall average. Of the three undisturbed study sites, Hasler Flats had the highest CV for Simpson index. 3.3.3 Regression analysis: assessing the relationship between vegetation and environment beta diversity Simple linear regression was used to test if environment Bw significantly predicted vegetation Bw both on the landslide and in the surrounding undisturbed terrain (Figure 3-2). 139 Figure 3-2. Landslide and undisturbed transect beta diversity (Bw) regression: vegetation vs environment. Blue symbols represent Cecil Lake transects, green symbols represent Beatton River transects, and orange symbols represent Hasler Flats transects. The blue dashed line is the regression line for landslide transect beta diversity (Bw). There was no significant regression for the undisturbed transects. A regression line with the R2 (coefficient of determination) value was drawn for the landslide transects, and a separate analysis was conducted for the undisturbed transects, to determine whether there was any relationship or correlation between vegetation beta diversity and environmental beta diversity. The fitted regression model was: Vegetation Bw = 1.617631918 + 3.525096525*(environment Bw). The regression for the landslide data was statistically significant (R2 = 0.75, F (1, 13) = 38.71, p = 0.000031). The standard error of the slope coefficient was 0.5767 indicating the observed values for the landslide data fell an 140 average of 0.58 units from the regression line. The low standard error suggested the model was precise enough to be used for predictions. Using the model, environment Bw significantly predicted vegetation Bw on landslides. The landslide transect vegetation beta diversity had a strong relationship with the beta diversity of the environmental variables. The R2 value of 0.7486 indicated that almost 75% of the variation in vegetation Bw was explained by environment Bw. The undisturbed transects did not yield a significant relationship between vegetation beta diversity and environment beta diversity. When the landslide and undisturbed plots were combined in a single analysis (not reported), there was a significant, though somewhat weaker relationship than for just the landslide plots. 3.4 Discussion The purpose of this research was to assess turnover of vegetation and site (environment) variables on landslides and to determine if there was any correlation between the two components. The research also investigated whether any correlations or relationships were influenced by size or age of the landslide disturbance, and whether there was a difference between landslide and undisturbed terrain areas. In this study, vegetation and environment beta diversity were generally higher on landslides than on the surrounding undisturbed terrain. In addition, there were notable differences in diversity between the three study areas of Beatton River, Cecil Lake, and Hasler Flats. Average transect vegetation and environment beta diversity were both higher on the landslide than on the surrounding undisturbed terrain for the Cecil Lake and Hasler Flats study areas, but for Beatton River, vegetation and environment beta diversity were higher on the undisturbed terrain. The lower beta diversity on the Beatton River landslide was likely because the slide was newer and more active than the other two study sites, resulting in large 141 areas of continuous bare ground. The Beatton River landslide also had the largest change in elevation from the top of the landslide to the bottom. In addition, this landslide was steeper and had more extreme exposures than the other two landslides, hindering the establishment of a variety of species across the disturbed area. The size of the three different study areas also likely affected the beta diversity values, as area influences both environmental gradients and the ecological mechanisms (such as species sorting) that drive spatial variation in species composition within a region unit (Heino et al. 2014). The occurrences of lower alpha diversity but higher beta diversity on the landslide compared to the surrounding terrain for some of the study areas may be because while mature forest species were lost, the patchiness of the landslide habitats enabled more novel assemblages of species or life history strategies (Boinot and Alignier 2023). The structural complexity and heterogeneity of pre-existing and remaining habitats contributes to local and regional biotic successional processes and ultimately influences beta diversity (Abbasi et al. 2023). This impact is mainly seen through changes in both taxonomic and functional beta diversity. At the local level, biotic processes are determined by species-energy relationships and availability of resources. Overall, vegetation beta diversity had a strong positive relationship with site or environment beta diversity on landslides in the study. As environment beta diversity increased, so did vegetation beta diversity. On the surrounding undisturbed terrain, however, there was no such relationship, either positive or negative. 142 3.4.1 Transect vegetation beta diversity 3.4.1.1 Vegetation beta diversity on the landslide There were notable differences in vegetation beta diversity between the three landslide study sites of Beatton River, Cecil Lake, and Hasler Flats. Average transect vegetation beta diversity (Bw) was highest on the Hasler Flats landslide, but Cecil Lake had the greatest range or variability. The higher variation of the landslide vegetation beta diversity for the Cecil Lake transects was likely directly due to the larger size of the landslide, its age, and the adjacent landscape. The passage of time allowed more types of vegetation to encroach from the surrounding landscape and to propagate from rafts and seed banks. Overall, the surrounding landscape makeup may have strongly influenced the recovery rate of species diversity (Furusawa et al. 2023). The higher among-site variability in plant species composition (beta diversity) on the Cecil Lake landslide suggests both a random, stochastic process of ecological drift and niche selection were critical for community assembly following the landslide disturbance. Field observations indicated the Cecil Lake landslide had a greater variety of site types in close proximity compared to Beatton River and Hasler Flats, allowing for the potential for higher turnover of plant species along any given transect distance. For example, the transect CEtl3c had the highest landslide vegetation beta diversity value (Bw = 5.5) and intersected a pond for ten metres of the transect length (Figure 3-3). This local site diversity likely contributed to the higher species turnover, as the pond and surrounding shorelines presented different habitats where new plants and unique communities could establish. Conversely, the Cecil Lake landslide transect with the lowest beta diversity, CEtl1 (for which Bw =1.9), had a 143 uniform site type consisting of dense shrubs throughout the transect (Figure 3-4). This uniformity is expressed in the lower turnover denoted by a lower Bw value. Figure 3-3. Cecil Lake landslide transect CEtl3c, showing high diversity of adjacent habitats as illustrated by Alnus spp. next to a pond, with sparsely vegetated south-facing slopes on the other side of the pond. (Photo August 25, 2018) Figure 3-4. Cecil Lake landslide transect CEtl1, showing lower diversity of adjacent habitats as illustrated by expanse of level ground and relatively homogeneous vegetation. (Photo August 25, 2017) 144 The higher average beta diversity of the Hasler Flats landslide reflects the abundance of very different site types situated very close to each other, coupled with the much smaller area of the Hasler Flats landslide (Heino et al. 2014), compared to the Cecil Lake landslide. The Hasler Flats landslide consisted of a series of interlocking young aspen cutblock rafts from a 15 year old clearcut stand, exposed horsts (ridges of sand or clay) and grabens, and pond site types. Following the landslide disturbance, vegetation dispersal distances would have been shorter both from the surrounding landscape and within the landslide area. In addition, small samples in areas with high gamma diversity tend to have inflated beta diversity values, due to the dependence on sample size that interacts with gamma diversity (Cao et al. 2021). Despite its high site type diversity, Hasler Flats likely showed the smallest range of transect vegetation beta diversity values because of its smaller sample size. Hasler Flats had less than half the number of landslide transects of Cecil Lake, so this could have reduced the range just because there were fewer samples to begin with. Although Hasler Flats had the highest mean vegetation beta diversity, it had the lowest CV. This lower CV indicates much less variation around the mean compared to Beatton River and Cecil Lake. This finding was in accord with field observations, as the Hasler Flats landslide overall appeared more consistent in diversity than Beatton River or Cecil Lake. The lower average vegetation beta diversity of the Beatton River landslide compared to the other two landslide study sites was likely due to the large proportion of the slide that was sparsely vegetated because of recent movement and steep, unstable slopes. The Beatton River landslide also contained the transect with the lowest overall vegetation beta diversity, BEtl4 (Figure 3-5), for which Bw = 0.8. The transect BEtl4 was on a sparsely vegetated 145 rubbly colluvial side slope and most of the vegetation was the invader exotic pioneers Melilotus officinalis and Melilotus albus (sweetclovers). Figure 3-5. Sweetclover (Melilotus spp.) carpeting the toe of the Beatton River landslide. Transect is BEtl4. (Photo August 16, 2017) Research has shown that sites with lower beta diversity may have higher alpha diversity, and the opposite can be true as well (Boinot and Alignier 2023). The background alpha diversity results of the present study showed variable outcomes. The Hasler Flats landslide had the highest richness (Figure 3-6), likely due to the many species rafted or otherwise dispersed from the adjacent aspen cutblock and the surrounding aspen (Populus tremuloides) and cottonwood (P. balsamifera ssp. balsamifera) stands with shrubby understories. The greater evenness, Shannon, and Simpson values at Cecil Lake may have resulted from the higher diversity of site types and the greater age and size of the landslide. Of the three landslides, 146 Beatton River exhibited the lowest vegetation richness, evenness, Shannon, and Simpson measures. The lower alpha diversity values for Beatton River were likely due to the younger age of the landslide, as well as the fact that it was still quite active, so even when plants did establish, they could quickly die off or slide downslope and be buried or otherwise damaged. Depending on the severity of the disturbance, landslides can cause a shift from stochastic or random to more deterministic or predictable plant community assembly, with distinct responses in terms of beta diversity (Seguin et al. 2013). More frequent or severe landslides can lead to a homogenised species distribution or a high proportion of invasive species, as found on the Beatton River transect BEtl1 (Figure 3-7), while a lower disturbance frequency or severity can cause greater stochasticity. Figure 3-6. Vegetation diversity on the Hasler Flats landslide, illustrated by dense layers of varied forbs, short shrubs, and tall shrubs. Transect is HAtl1a. (Photo July 28, 2018) 147 Figure 3-7. Invasive/exotic plants Lactuca serriola (prickly lettuce), Sonchus spp., Melilotus spp., and Elymus repens (quackgrass) on the Beatton River landslide. Transect shown is BEtl1. (Photo August 17, 2017) 3.4.1.2 Vegetation beta diversity in the surrounding undisturbed terrain The variable levels of beta diversity in the undisturbed areas surrounding each landslide highlights the complexities of assessing beta diversity. The greater range of transect vegetation beta diversity values and the higher variability in the adjacent undisturbed terrain of the Beatton River site compared to Cecil Lake and Hasler Flats may have been due to the higher diversity of ecological habitats surrounding the Beatton River landslide. Habitats included mature conifer forest, mature aspen forest, mature cottonwood forest (Figure 3-8), shrublands, and grasslands. 148 Figure 3-8. Example of moist cottonwood (Populus balsamifera ssp. balsamifera) habitat in undisturbed terrain at Beatton River. Transect is BEtu1a. (Photo August 26, 2018) Cecil Lake exhibited the lowest range of vegetation beta diversity values of the three undisturbed study sites but yielded the highest average undisturbed vegetation transect beta diversity. The Cecil Lake undisturbed study site also had the lowest CV, indicating less variation around the mean compared to Beatton River and Hasler Flats. The comparatively 149 higher undisturbed transect vegetation beta diversity for Cecil Lake could have been due to the larger area, increasing the likelihood of transects falling on more heterogeneous terrain. Compared to the associated landslides, the mean vegetation beta diversity was lower in the surrounding undisturbed terrain for Cecil Lake and Hasler Flats, but higher in the undisturbed terrain than on the landslide at the Beatton River study area. Although there may be no significant differences in species diversity between an undisturbed and landslide community, much higher among-site variability may be found on landslides (Furusawa et al. 2023). The generally lower vegetation beta diversity on the undisturbed terrain for the three study areas was likely due to the presence of greater areas of homogeneous habitats, such as the adjacent cutblock next to Hasler Flats (Figure 3-9), as well as the passage of time that allowed certain successional species to outcompete pioneers and take over. Figure 3-9. Relatively homogeneous vegetation (young aspen – Populus tremuloides) in undisturbed terrain at Hasler Flats. Transect is HAtu1a. (Photo August 10, 2018) 150 Overall, the assessment of vegetation alpha diversity in the undisturbed terrain among the three study areas revealed variability between study sites, with higher alpha diversity values for Beatton River and Hasler Flats and lower values for Cecil Lake in the undisturbed terrain compared to the landslide areas. The higher alpha diversity richness value of Hasler Flats compared to Cecil Lake may have been due to the greater array of shrub species in the terrain surrounding the Hasler Flats landslide. Hasler Flats also yielded the highest Shannon and Simpson values, further suggesting the surrounding undisturbed terrain vegetation had higher alpha diversity than the Beatton River and Cecil Lake study sites. Cecil Lake had the lowest Shannon and Simpson values, indicating the vegetation on the undisturbed terrain was less diverse. The similar evenness for all three undisturbed sites suggests the sites were similar in the proportional distribution of various species. It should be noted that “undisturbed” was a relative term, and for all three landslides there were signs of past disturbance in the surrounding terrain. There was evidence of older landslides, historical logging, cultivation of fields, and possibly other natural disturbances. All these factors undoubtedly contributed to a diversity of plant community growth forms and successional stages. 3.4.2 Transect environment beta diversity 3.4.2.1 Environment beta diversity on the landslide Overall, the environment beta diversity values on the landslides were much lower than vegetation beta diversity and had smaller ranges, suggesting a lower turnover of site diversity compared to vegetation. Disturbance such as a landslide can either increase or decrease environmental heterogeneity, and likewise either increase or decrease beta diversity (Maab et al. 2014). The environment of an area can locally filter dispersing plant species, which may 151 establish either through niche partitioning or dispersal (Maab et al. 2014). The variable environment beta diversity values among landslides in this study indicate different drivers influencing diversity for each landslide. Just as for vegetation beta diversity, the Cecil Lake landslide study site had the greatest range of environment beta diversity values, while Hasler Flats showed the smallest range. The higher range of beta diversity values at Cecil Lake could have been due to the greater diversity of site types observed on the landslide compared to Hasler Flats. The findings could also simply be explained by the fact that Cecil Lake landslide was almost 40 times larger and had more transects than Hasler Flats, increasing the potential for a random transect to fall on a diverse site. However, Hasler Flats had a much higher sampling intensity given its size, so the influence may not have been that great. Either way, the higher range indicates an increased variability in turnover of environment features on the Cecil Lake landslide. The Cecil Lake and Hasler Flats landslide study sites were tied for the highest environment beta diversity of the three landslide study sites, even though Cecil Lake landslide was more variable. Hasler Flats was a smaller and younger slide, and it appears these factors contributed to a higher beta diversity. The landslide was still in the process of weathering and becoming more subdued. The lower average environment beta diversity of Beatton River is consistent with field observations, as large parts of the landslide were uniform in slope, aspect, and substrate. However, Beatton River also had the highest CV (over 100%), indicating a large amount of variation among transects. The variability in environment alpha diversity results for the three landslide study sites highlights the influence of space and time on environment heterogeneity (Maab et al. 2014). The higher landslide environment alpha diversity results for Beatton River contrasted with the lower beta diversity results. Higher alpha diversity could be explained by the site 152 diversity present along some of the Beatton River transects. One of the transects, BEtl6, passed through rubbly boulders, across a pond, and then up a steep slope (Figure 3-10). Other transects on the landslide had patches with no vegetation. The lower environment richness of Cecil Lake compared to Beatton River and Hasler Flats could be due to the age of the landslide, as geomorphological features weather over time, becoming more uniform. The lower alpha diversity values for Hasler Flats may have been a result of the small size of the landslide and the more consistent configuration of the environment features. Figure 3-10. Beatton River landslide transect BEtl6, showing microsite diversity of hummocks, depressions, ponds, and fissures. (Photo taken September 5, 2018) 153 3.4.2.2 Environment beta diversity in the surrounding undisturbed terrain Overall, mean environment beta diversity values for the undisturbed transects were lower than for the landslide transects, but on an individual basis the Cecil Lake and Hasler Flats undisturbed study sites had lower average beta diversity while Beatton River had higher beta diversity compared to the associated landslides. Lower environment beta diversity in the undisturbed terrain could be due to the surfaces exhibiting a lower diversity of microsites over short distances compared to the landslides, as a result of minimal disturbance over time. The higher average beta diversity value for Hasler Flats and the lowest average beta diversity for Cecil Lake undisturbed samples could be explained by the fact that Hasler Flats samples were in more broken terrain (Figure 3-11) in the surrounding area, while Cecil Lake undisturbed samples fell in more uniform terrain (Figure 3-12). The similarity in beta diversity range for Beatton River and Hasler Flats is likely due to the similar undisturbed terrain of these two study sites. 154 Figure 3-11. Hasler Flats undisturbed transect HAtu3 showing broken terrain. Transect traverses uneven terrain and small fissures for the first 15 m (foreground), then ascends a steep slope for 8 m (visible in the distance), and finally levels out (evident in the photo where the sky shows through). (Photo August 3, 2018) 155 Figure 3-12. Cecil Lake undisturbed transect CEtu2 showing uniform, level terrain. (Photo August 17, 2018) The variable environment alpha diversity of transects on undisturbed sites was reflective of the variation among landslide study areas. The higher environment alpha diversity values of richness, evenness, Shannon, and Simpson values for transects in the Beatton River undisturbed terrain could be explained by the fact that Beatton River had a variety of site types and ecosystems. Conversely, Hasler Flats exhibited the lowest richness, while Cecil Lake had the lowest evenness, Shannon, and Simpson values. The lower environment 156 richness of the undisturbed Hasler Flats transects reflects more uniform terrain and may also be due to the smaller area over which transect locations were selected. The lower alpha diversity values for Cecil Lake may have been the result of a more subdued landscape overall, given the larger scale of the site as well as its advanced age. 3.4.3 Relationship between transect vegetation and environment beta diversity Environmental factors shape both local and regional biotic processes, mainly through changes in functional and taxonomic beta diversity patterns (Abbasi et al. 2023) but also alpha diversity patterns (Boinot and Alignier 2023). The current study in the Peace River Region revealed a significant relationship between vegetation beta diversity and environment beta diversity on the landslide study sites, while there was no such relationship on the undisturbed sites. Generally, as environment beta diversity increased on the landslides, so did vegetation beta diversity, even though the environment beta diversity values were notably lower than those for vegetation beta diversity. The stronger association between vegetation and environment beta diversity on the Cecil Lake landslide study site compared to Beatton River and Hasler Flats could be explained by the greater age of this landslide, where plant communities have evolved with the changing microsites over time. It is also possible the plant communities altered the environment as time passed. The lack of a strong relationship between environment and vegetation beta diversity on the undisturbed areas is most likely due to the overall lower beta diversity for both environment and vegetation on these sites. The sites have stabilised, and thus more resilient plant species have tended to dominate. Research has shown beta diversity is positively associated with local topographical heterogeneity as well as with community-level niche specialisation and niche marginality (Cao et al. 2021). Higher niche marginality, or ability to occupy the peripheries, indicates 157 larger niche space, which may allow more species to use more variable resources, while higher niche specialisation enables species to specialise on narrower subsets of the resources present. Therefore, on a highly variable topography where niche specialisation or marginalisation are at work, a greater turnover of species can be found from one point to another across a given distance. Local processes of topographical heterogeneity and the resultant niche differentiation drive beta diversity at scales of 15-52 ha. Greater environmental variability on a site may ultimately lead to reduced evenness in communities (Furusawa et al. 2023). Connectivity between habitats is important for driving the recovery of disturbed sites such as landslides. Connectivity is a product of the synergies between natural history, dispersal mechanisms, and the quantity, quality, and pattern of habitat patches, at the landscape level (Chiantore et al. 2018). Connectedness of habitats and the successional state of ecological communities together play an important role in understanding consequences of change in different ecosystems. Of the three landslides studied, Hasler Flats appeared to have the greatest connectivity, mainly due to its small size and the presence of many rafts. From the perspective of beta diversity as an indicator of ecological connectivity, recovery does not necessarily depend on the available species at the regional level. Instead, local ecological attributes affiliated with beta diversity and the creation of habitats by living organisms drive community assembly by way of species replacement (turnover) and habitat filtering in disturbed areas. Habitats and communities can become homogenised with an increased disturbance frequency or severity, exhibiting an accelerated homogenisation with increasing scale of disturbance. This condition appeared to be present on portions of the Beatton River landslide, where disturbance was still active and exotics dominated large portions of the 158 landslide. Boinot and Alignier (2023) found alpha and beta diversity can be maintained with a variety of weed species and life strategies in highly disturbed sites, even if rare and more sensitive species are removed from the community. Homogeneous sites also recover faster, which results in diminished complexity and biodiversity over time (Chiantore et al. 2018). 3.4.4 Limitations One of the limitations to the study was the fact that the number of samples and sampling intensity per hectare at each study site were not the same. The only study area with the same amount of transects on the landslide and in the surrounding undisturbed terrain was Hasler Flats. Another limitation was that not all transects were of equal length or had the same number of plots in a sample. For example, two transects could have both been 30 m long, but one might have only 20 plots because a portion of the transect was skipped once there were no new species. Plots were sampled again along the transect if there was a new type, but it is possible there were microsites in between that contained new species. Therefore, beta diversity could have been underestimated. Further, the landslides varied in age, type, and size, rendering the results more challenging to generalise. Overall, however, the methods used did provide a useful synopsis of beta diversity on landslides and in the surrounding terrain, and the study provided many insights into turnover on diverse disturbed sites. 3.5 Conclusions and Recommendations Although there was some variability between transects on some landslides, overall, average vegetation and environment beta diversity (Bw) appears to be higher on landslides than on surrounding undisturbed terrain for sites in northeastern British Columbia. The trend is more pronounced for vegetation beta diversity. There is a strong relationship between vegetation 159 beta diversity and environment beta diversity on the landslides, even though the environment beta diversity values are much smaller. This finding suggests that environment beta diversity does not need to be very high to produce increased vegetation beta diversity. The differences in vegetation and environment beta diversity values between landslides of differing sizes and ages indicates that spatial and temporal factors may influence the level of turnover on a given landslide. The smallest landslide (Hasler Flats) had the highest beta diversity, and it was the second oldest of three landslides. The newest and most active landslide (Beatton River) had the lowest beta diversity. The findings of this study of beta diversity on landslides can help inform restoration planning for other landslides in the Peace River Region, as well as for low-gradient, deep-seated landslides throughout the circumboreal zone which occur on glaciolacustrine unconsolidated material. Beta diversity is a primary signal of the health of a community and the proper functioning of ecosystems. It can provide important information about ecosystem mechanisms for the purposes of restoration. However, patterns of beta diversity on their own should not be used as a benchmark for restoration, since observed change in beta diversity in disturbed areas such as landslides is chiefly due to random sampling effects influenced by changes in local community size (Myers et al. 2015). Sites with very similar habitat might have quite different makeup of species because of the order that species arrive on the landslide. This phenomenon can produce multiple stable equilibria. Knowledge about the connections between ecosystem disturbance and taxonomic, functional, and beta diversity of landslides may provide crucial information for natural resources management (De et al. 2023) as well as restoration. Beta diversity presents important 160 potential for management of environmental monitoring and conservation since diverse processes can result in the same beta diversity pattern (Maab et al. 2014). The present study of landslides has shown that when only a few plant species prevail because of accelerated habitat disruption, the species composition becomes more uniform and the diversity of the community decreases, as evident on the Beatton River landslide. High alpha diversity may obscure deleterious effects of habitat disturbances on plant species diversity (Dehling and Dehling 2023). Just as for beta diversity, alpha diversity alone should not be viewed as a representative indicator of disturbance impacts since it does not consider possible local or regional changes in species composition or turnover. Habitat alterations resulting from disturbances such as landslides can eliminate vital functional roles, reducing an ecologically complex system to one that is simpler. In this situation, unique native species disappear, and more common and often invasive species take over, creating homogenised communities that diminish regional diversity. Alpha and beta diversity can be maintained with a variety of weed species and life history strategies in simple, severely disturbed areas, even when uncommon and more disturbance-sensitive species are eliminated (Boinot and Alignier 2023). However, this type of diversity is not a good indicator for conservation or healthy functioning of ecosystems, because the original natural community system has been modified. As there are different drivers for alpha and beta diversity, both types of diversity can be advanced by implementing different arrays of management practices (Boinot and Alignier 2023). Any attempts at restoration of landslides should retain patchiness and maintain beta diversity. As climates change in an unpredictable way in coming years, beta diversity may provide important wildlife habitat at various scales and enhance resilience of populations in uncertain times. 161 3.6 References Abbasi, U.A., E. Mattsson, S.P. Nissanka, and A. Ali. 2023. Species α-diversity promotes but β-diversity restricts aboveground biomass in tropical forests, depending on stand structure and environmental factors. Journal of Forest Research, 34: 889-901. Anderson, M. J., T.O. Crist, J.M. Chase, M. Vellend, B.D. Inouye, A.L. Freestone, N.J. Sanders, H.V. Cornell, L.S. Comita, K.F. Davies, S.P. Harrison, N.J.B. Kraft, J.C. Stegen, and N.G. Swenson. 2011. Navigating the multiple meanings of β diversity: a roadmap for the practicing ecologist. Ecology Letters 14, 19–28. doi:10.1111/j.1461-0248.2010.01552.x Barwell, L.J., N.J.B. Isaac, and W.E. Kunin. 2015. Measuring β-diversity with species abundance data. Journal of Animal Ecology, 84: 1112-1122. BC Gov (British Columbia Ministry of Forests and Range [BCMFR] and British Columbia Ministry of Environment [BCMOE]). 2010. Field manual for describing terrestrial ecosystems. 2nd edn. BCMFR Research Branch and BCMOE Resource Inventory Branch, Victoria, B.C. (Reprint with updates 2015.) Boinot, S., and A. Alignier. 2023. Discrepancies between the drivers of alpha and beta plant diversity in arable field margins. Proceedings Royal Society B, 290: 20222179 (11 pages). Bray, J.R., and J.T. Curtis. 1957. An ordination of the upland forest communities in Southern Wisconsin. Ecological Monographs, 27:325-349. Cao, K. et al. 2021. Species packing and the latitudinal gradient in beta diversity. Proceedings Royal Society B, 288. 20203045 (8 pages). Chao, A., and C.-H. Chiu. 2016. Bridging the variance and diversity decomposition approaches to beta diversity via similarity and differentiation measures. Methods in Ecology and Evolution, 7: 919-928. Chiantore, M., S.F. Thrush, V. Asnaghi, and J.E. Hewitt. 2018. The multiple roles of βdiversity help untangle community assembly processes affecting recovery of temperate rocky shores. Royal Society Open Science, 5: 171700 (9 pages). De, K., A.P. Singh, A.S. Sarkar, K. Singh, M. Siliwal, V.P. Uniyal, and S.A. Hussain. 2023. Relationship between species richness, taxonomic distinctness, functional diversity, and local contribution to β diversity and effects of habitat disturbance in the riparian spider community of the Ganga River, India. Ecological Processes, 12: 13 (13 pages). Dehling, D.M., and J.M. Dehling. 2023. Elevated alpha diversity in disturbed sites obscures regional decline and homogenization of amphibian taxonomic, functional, and phylogenetic diversity. Scientific Reports, 13: 1710 (9 pages). 162 Furusawa, J., K. Makoto, and S. Utsumi. 2023. A large-scale field experiment of artificially caused landslides with replications revealed the response of the ground-dwelling beetle community to landslides. Ecology and Evolution, 13: e9939. https://doi.org/10.1002/ece3.9939 Heino, J., A.S. Melo, and L.M. Bini. 2015. Reconceptualising the beta diversityenvironmental heterogeneity relationship in running water systems. Freshwater Biology, 60: 223-235. Jaccard, P. 1908. Nouvelle recherches sur la distribution florale. Bulletin de la Societe Vaudoise des Sciences Naturelles, 44: 223-270. Jurasinski, G., V. Retzer, and C. Beierkuhnlein. 2009. Inventory, differentiation, and proportional diversity: a consistent terminology for quantifying species diversity. Oecologia, 159:15- 26 DOI 10.1007/s00442-008-1190-z Koleff, P., K. Gaston, and J. Lennon. 2003. Measuring beta diversity for presence–absence data. Journal of Animal Ecology, 72: 367-382. Lennon, J., P. Koleff, J.J.D. Greenwood, and K.J. Gaston. The geographical structure of British bird distributions: diversity, spatial turnover, and scale. Journal of Animal Ecology, 70: 966-979. Maab, S., M. Migliorini, M. Rillig, and T. Caruso. 2014. Disturbance, neutral theory, and patterns of beta diversity in soil communities. Ecology and Evolution, 4(24): 4766-4774. Magurran, A. 2004. Measuring biological diversity. Blackwell Publishing. Oxford, UK. 256 p. McCune, B., and D. Keon. 2002. Equations for potential annual direct incident radiation and heat load. Journal of Vegetation Science, 13: 603-606. McCune, B., and M. J. Mefford. 2016. PC-ORD. Multivariate Analysis of Ecological Data. Version 7. MjM Software Design, Gleneden Beach, Oregon, U.S.A. McCune, B., and M. J. Mefford. 2018. PC-ORD. Multivariate Analysis of Ecological Data. Version 7.10. MjM Software Design, Gleneden Beach, Oregon, U.S.A. Moreno, C., and P. Rodriguez. 2010. A consistent terminology for quantifying species diversity? Oecologia, 163:279-282. Myers, J.A., J.M. Chase, R.M. Crandall, and I. Jimenez. 2015. Disturbance alters betadiversity but not the relative importance of community assembly mechanisms. Journal of Ecology, 103: 1291-1299. 163 Seguin, A., D. Gravel, and P. Archambault. 2014. Effect of disturbance regime on alpha and beta diversity of rock pools. Diversity, 6: 1-17. Sørensen, T. 1948. A method of establishing groups of equal amplitude in plant sociology based on similarity of species content. Biologiske Skrifter, 5:1-34. Tuomisto, H. 2010a. A diversity of beta diversities: straightening up a concept gone awry. Part1. Defining beta diversity as a function of alpha and gamma diversity. Ecography, 33:222. Tuomisto, H. 2010b. A diversity of beta diversities: straightening up a concept gone awry. Part 2. Quantifying beta diversity and related phenomena. Ecography, 33: 23-45. Tuomisto, H. 2010c. A consistent terminology for quantifying species diversity? Yes, it does exist. Oecologia, 164:853-860. Whittaker, R. 1960. Vegetation of the Siskiyou Mountains, Oregon and California. Ecological Monographs, 30: 279-338. Whittaker, R. 1972. Evolution and measurement of species diversity. Taxon, 21(2/3): 213251. Wilson, M.V., and A. Shmida. 1984. Measuring beta diversity with presence-absence data. Journal of Ecology, 72(3) (Nov. 1984): 1055-1064. 164 Chapter 4. Landslide-generated ponds in the Fort St. John area, British Columbia: Characteristics, distribution, and ecological implications 4.1. Introduction Traditionally, landslides have been viewed as destructive to the habitats and ecology of a landscape, in that they remove vegetation, bury other vegetation, disintegrate soil profiles, fill in water bodies, and alter hydrology. However, the same processes that are seen as destructive can serve to enhance ecological diversity by creating new geomorphic and hydrological features. In addition to depositing sediment nutrients and habitat debris into river systems and changing stream morphology, landslides often produce a diverse array of topographies on the landslide surface, with many pond depressions that contain water ephemerally, seasonally, or year-round. The size, number, and distribution of such water bodies is variable and may depend on the material of the landslide as well as the landslide type. The focus of the research reported here is the ecological contribution of these landslide ponds to diversity and implications for management. Water bodies on landslides form important habitats that contribute to biodiversity in many different environments (Geertsema and Pojar 2007; Shapley et al. 2019). At the landscape level, these aquatic habitats can create a network of reproductive and dispersal routes for pond-dependent fauna such as invertebrates, amphibians, and reptiles; this network of various sizes of ponds provides more value than one large lake (Pop and Chitu 2013). While landslides themselves create ponds, they also can provide ideal habitat for beavers (Castor canadensis) (Krueger and Johnson 2016), and beavers can then further modify the geomorphology and hydrology of the landslide (Butler and Malanson, 1995; Westbrook et al. 2017) and increase the ecological diversity of ponds (Nummi et al. 2019). 165 Although few detailed studies of landslide ponds have been reported, existing research has revealed interesting characteristics. Sasaki and Sugai (2015) found that landslide ponds in the mountains of Japan chiefly occur on bigger landslides, in large or deep depressions along scarps, or in small depressions at pressure ridges, and are mainly fed by groundwater. The authors of the study also found that pond size is constrained by the topography of the landslide, and ponds at different stages of development can be present concurrently due to new activity on an existing landslide. Ponds can persist and expand over time (Cruden et al. 1997). Surface water infiltration through boulder debris and fractures on an existing slide can saturate the substrate and facilitate further slide activity (Coe et al. 2003), which then forms new depressions that develop into ponds. On larger slides, different temperature and water regimes may exist among head, body, and toe positions of the landslide due to differences in elevation, which can influence infiltration and pond persistence (Coe et al. 2003). Coe et al. (2009) found that pond location and persistence may be controlled by basal topography, in that the landslide ponds may persist in one location for over 100 years even while the landslide material moves around, below, and through the pond. The studies discussed above have helped to explain pond distribution and development on landslides in other parts of the world. The key purposes of the research reported here are to compile an inventory of landslide ponds in a designated study area of the Peace River Region of northeastern British Columbia, Canada using GIS applications, to describe characteristics of size and distribution of these ponds within landslides and between landslide types, to identify ecological importance, and to provide recommendations for management. The study of landslides in the Charlie Lake 1:250,000 mapsheet area and in the Peace River Region in general is of interest to various government and industrial entities because of the 166 potential and existing impacts of mass movement events on industrial development and public safety. Some of the earliest geomorphology studies were carried out in anticipation of a proposed hydroelectric dam, Site C (now under construction), near the town of Taylor (Mathews 1978; Catto 1991). These investigations focused on the geological setting of historic landslides and attempted to determine slide mechanics. Since then, major work has been done to describe the geological history and stratigraphic makeup of the mapsheet area (Mathews 1978, 1980; Bobrowsky et al. 1990; Bobrowsky and Rutter 1992; Catto et al. 1996; Hartman 2005). In recent years, more detailed assessments of movement mechanisms have been performed on specific landslides, such as the Cache Creek slide (Van Esch 2012), the Attachie slide (Fletcher et al. 2002; Van Esch 2012), and the Beatton River slide (Dandurand 2018). An inventory of all landslides in the Charlie Lake mapsheet area was compiled by Severin (2004), with each mapped landslide categorised according to factors such as movement type and activity level. Severin (2004) found that the majority of identified landslides occurred in the Peace River and Beatton River Valleys within the pre-glacial valley limits, where valley fill and rebound features in the shale bedrock are more prominent. In the current research, the aim is to investigate landslides in the Peace River Region with a focus on landslide ponds and their potential ecological value, using the same study area as Severin (2004). The objectives of the landslide pond research were to: 1) Obtain a snapshot in time of pond locations on landslides in the study area; 2) Enumerate and graphically illustrate pond area and distribution in relation to landslide type and geomorphic location; and 3) Investigate potential ecological and management implications of the findings. 167 What follows is a synthesis of the research and findings of the landslide pond study. First, the study area is described, followed by a detailed description of the methods that were employed to address the research question regarding the nature of landslide pond presence, abundance, distribution, and possible ecological roles in a specified area of the Peace River Region. The results are presented, including some representative pond photographs from a few pond-bearing landslides in the area. Presentation of the results is followed by a discussion of key findings regarding ponds on landslides and possible ecological implications. The discussion includes a section on the influence of beavers, which often play a role of keystone species at the landscape level. Finally, the chapter concludes with some recommendations for further research and land management considerations. The work reported here is the first known large-scale detailed regional inventory and description of ponds on landslides. 4.2. Study area The study area covers the entire NTS (National Topographic System) mapsheet 94A (Charlie Lake mapsheet, 1:250,000), an area of approximately 16,000 km2 (1,600,000 ha) that includes the communities of Fort St. John, Hudson’s Hope, and Taylor in the Peace River Region of British Columbia (Figure 4-1). The Beatton River and Cecil Lake Landslide study sites described in Chapters 2 and 3 are located within this mapsheet. The area is represented by the Boreal White and Black Spruce (BWBS) biogeoclimatic zone (Meidinger and Pojar 1991) and is completely within the boreal forest biome. The climate is temperate, with warm, wet summers and cold winters (Meidinger and Pojar 1991). Much of the precipitation falls in the summer months. Vegetation consists of pure and mixed conifer and aspen forests on the uplands, transitional aspen parkland, and grasslands along the river slopes. 168 Historically, the main forms of land use in the area were farming, forestry, and fur trapping. However, in recent decades, oil and gas activity has expanded exponentially, greatly increasing the industrial footprint on the land through exploration, extraction, and infrastructure. Figure 4-1. Landslide ponds study area – Mapsheet 94A (1:250,000) Peace River Region of northeastern British Columbia, Canada. The yellow areas indicate history of landslides along the Peace River and its tributaries, based on shape file linework by Severin (2004). Shape file linework imagery reproduced with permission. 169 The geology of the area consists of Quaternary stratigraphy from the historical episodes of Glacial Lake Mathews and Glacial Lake Peace (Mathews 1978, 1980) formed by successive advances and retreats of Cordilleran and Laurentide ice sheets. Many of the sediments are fine clay and silt overlaying shale bedrock (Catto 1991). The Shaftesbury (marine shale) and Dunvegan (marine and deltaic sandstone with shale interbeds) formations prevail in this area (Mathews 1978), and most existing landslides occurred within the Lower Cretaceous Shaftesbury Formation (Severin 2004). Many tributaries of the Peace River were formed by the rapid carving of valleys following isostatic rebound as the glaciers retreated and the land masses rose (Mathews 1978; Catto 1991; Hartman and Clague 2008). Prehistoric and contemporary landslides are abundant along the Peace River and its tributaries. 4.3. Methods This study used computerised geographic information system (GIS) techniques and large Excel spreadsheets to compile and analyse the distribution and spatial characteristics of pond-bearing landslides and their associated water bodies within the area of interest. It also incorporated some field observations obtained during data collection for Chapters 2 and 3 to enhance the discussion on pond description and development. To limit scope for the time-intensive digitising and data entry work, the study was restricted to a specific 1:250,000 mapsheet of the Peace River Region, mapsheet 94A, and only ponds on landslides were inventoried and described on this 16,000 km2 area. Ponds were digitised by hand rather than using digital image processing (i.e. supervised image analysis), as many of the ponds were smaller than what could be captured in processing, and the aim was to map all ponds visually discernible on the imagery. 170 To begin the mapping of landslide ponds, Mapsheet 94A (NTS 2021) was loaded from the World Imagery layer (World Imagery 2021) into the Global Mapper GIS application (Global Mapper 2021). Google Earth Pro imagery (Google Earth Pro 2021) was loaded on a separate computer monitor. World Imagery and Google Earth Pro are platforms that synthesise aerial photographs, and it was these photographs that were analysed. The resolution of both sets of imagery was intermediate, with World Imagery having a slightly better resolution. The two sets of imagery were used as cross-references to each other, to verify whether the features identified were indeed landslides and ponds, or something else. While the World Imagery had clearer features, Google Earth Pro had the advantage of providing a 3D view as well as rapid zooming capabilities. The vintage for both types of imagery varied and covered a range of years, typically 2006 to 2018. Using the World Imagery map layer and a map tile grid in Global Mapper, and Google Earth Pro as a cross-reference, the entire mapsheet was systematically assessed visually for the presence of ponds, and all ponds located on landslides were digitised working at a scale of approximately 1:500 (Figure 4-2). The ponds were then labelled with numbers and classified as either ponds, dried out ponds, or wetlands. Each pond was assigned a unique number. Although some ponds were partially or completely dried out, all were counted, since the drying appeared to be seasonal. Early spring imagery was not always available to confirm the extent of water pooling, but other indicators such as variations in vegetation were used to verify. The occurrence and extent of dried ponds was evidenced by aquatic or wetland vegetation still present, mainly recognisable as cattail (Typha spp. – most likely Typha latifolia), which contrasted in texture and colour against the surrounding terrain. Because 171 wetlands in general are essentially a type of pond, they were combined with the ponds when compiling and analysing the data. Figure 4-2. Sample of landslide pond mapping. Location is Cecil Lake. Ponds are outlined in blue. Yellow lines indicate landslide features that had been previously mapped by Severin (2004). Severin landslide shape file lines used with permission. Once the ponds were mapped, Severin’s (2004) landslide shape file was loaded into Global Mapper as an overlay on the pond polygons. Severin’s digital GIS rendering of landslides (2004) was in the format of linework rather than enclosed area polygons (see Figure 4-2), and thus individual areas of these landslides were not obtainable in the GIS dataset. Therefore, the table from the inventory created by Severin and the landslide areas provided in Appendix IV of his thesis were used to obtain the initial total area of landslides. In the present study, a small number of additional landslides that were not included in the 2004 study were identified and digitised, as well as portions of other landslides where new movement had 172 occurred since the 2004 work done by Severin. The new area was added to the total landslide area prior to analysis of landslide ponds. No minimum or maximum size limit was assigned for either ponds or landslides; anything that could be discerned by zooming in on the imagery was mapped. A large Excel spreadsheet was created to record various data related to the ponds and associated landslides (Appendix 7). In the spreadsheet, a unique number was assigned to each pond-bearing landslide, in numerical order starting from the first landslide with ponds encountered in the pond digitising exercise. The landslides were not numerically labelled in Global Mapper, as several landslides were either eliminated or renumbered in the final spreadsheet list. However, the geographical location (UTMs – Universal Transverse Mercator) of the centre of each pond-bearing landslide was recorded in the spreadsheet. Once pond and landslide numbers were assigned, the general location of each pond on the landslide was recorded in the spreadsheet, for example if it was above or below a scarp or on a debris apron. The geomorphic location of each pond was also recorded and classified as to whether it was on the head, body, or toe of the landslide, using a simplified version of Cruden and Varne’s (1996) rendering of landslide anatomy (Figure 4-3). In some situations, ponds appeared to be on the borderline between geomorphic locations. If a pond seemed borderline between the head and the body, the pond was classified as being on the head if it was at the base of the main scarp before the slope changed by more than approximately 5 degrees. If the pond was located after a change in slope, it was classified as being on the body. A similar method was applied to ponds that appeared borderline between the body and the toe, in that ponds located at the base of the body before the slope changed significantly were classified as being on the body, and those occurring after a slope change were classified 173 as being on the toe. The Global Mapper Profile tool applied on an Advanced Spaceborne Thermal Emission and Reflection Radiometer (ASTER) Global Digital Elevation Model (DEM) image of 1 arc-second resolution (ASTER GDEM v2 2021) was used to assist in determining slope. Geomorphic location classifications were assigned initially by incorporating information on landslides found in Cruden and Varnes (1996) and Highland and Bobrowsky (2008) but decisions were ultimately subjective based on the characteristics of each individual location. Figure 4-3. Simplified anatomy of a landslide showing head, body, and toe. Adapted from Cruden and Varnes 1996. Credit: Transportation Research Board. 1996. Landslides: Investigations and Mitigation. Special Report 247. https://doi.org/10.17226/11057. Reproduced with permission from the National Academy of Sciences, Courtesy of the National Academies Press, Washington, D.C. For each pond, the size in km2 was recorded as calculated in Global Mapper, and values were then converted to hectares (ha) for the purposes of the study. The movement type of each pond-bearing landslide was classified according to the relevant descriptions and diagrams 174 laid out in Severin’s (2004) thesis. In the study area, six pond-bearing landslide types were identified: compound failure, mobile flow, multi-level rotational failure, retrogressive rotational failure, rotational failure, and shallow retrogressive rotational failure. The definitions of these types are outlined in Severin (2004) and are based on key landslide classification works (Varnes 1978; Hutchinson 1988; Cruden and Varnes 1996). However, Severin modified some classifications to reflect the unique characteristics of landslides in the Peace River Region. Compound failures occur on pre-sheared surfaces in bentonitic-rich layers of Shaftesbury shale and generally have a low angle. As the slope range parameter of compound failures was not specified in Severin (2004), a subjective judgment was made in the present study to classify as a compound failure any slide with a relatively long, straight profile and an average slope of approximately 20% or less. This classification method relied heavily on the Global Mapper Profile feature to assess slope profile characteristics. Mobile flows usually occur in gullies along the river and fan out at the bottom of the gully. They are shallow and are triggered during heavy rainfall and snowmelt seasons. Rotational failures develop along pre-sheared surfaces mainly in pre-glacial sediments, mostly on south-facing slopes. The debris is often broken up across the slope, rather than appearing as a classic slumped block. Multi-level rotational failures take place along several weak shear planes at different levels. The failures can occur either separately or simultaneously. Retrogressive rotational failures develop when rupture surfaces lengthen along a weak layer in the slope and there is progression of a single rotational slip upslope, ending at a curved back scarp. The initial small failure destabilises the toe, causing further slumping, and there are often back-tilted blocks of debris present. Shallow retrogressive rotational failures are similar to retrogressive rotational failures, but only occur in Glacial Lake Peace clay. The till below acts as a base that the failure plane cannot penetrate. The basal shear plane normally 175 coincides with the surface of the Wisconsinan till. The sediment remobilises and flows over a stable bench and quickly disintegrates at a slope angle of 3.5 degrees. Although the original landslide inventory (Severin 2004) categorised each landslide by movement type and assigned it a reference number, no digital or hard copy geographical reference database or information in the GIS shape files was available to determine the reference number or landslide type designation of any specific landslide within the mapsheet study area. Therefore, classification of landslide types in the current study mostly relied on Severin’s geomorphological symbols in the map file and descriptions in the thesis text, combined with the Profile feature in Global Mapper applied on a Digital Elevation Model (DEM) image and my understanding of geomorphological features. The activity level of each slide was classified using Severin’s (2004) activity classifications, as that information was present in the shape files themselves. Severin’s classifications for activity level included Very Active, Active, Low Activity – Dormant, Low Activity – Abandoned, and Modified. Low Activity – Dormant landslides refer to those landslides that are probably older than 50 years, but still have active erosion near the toe, while Low Activity – Abandoned landslides are older than 50 years and do not currently have active erosion at the toe. Landslides were classified as Active that were originally Dormant if there was any new activity on them, as Active slides are defined as those with activity in the last 50 years (Severin 2004). Modified landslides were those that had been stabilised with artificial earth works following failure. The last column in the spreadsheet was a comments section, where pertinent information about the ponds or landslides was recorded. After the initial inventory was completed, each map tile of the mapsheet was methodically re-checked to verify classifications and record any 176 missed features. Some pond polygons were deleted from the final list due to not falling on an actual landslide, while other pond polygons were deleted because they were not actually ponds, but instead were shadows. The ponds that remained in the data set retained their original number designations. Therefore, there were some gaps in the final numbering. For the final spreadsheet, comments were modified or deleted as issues were addressed. This spreadsheet was then used as a baseline for various calculations and graphs. The overall total number and area of ponds and the total number and area of landslides for the mapsheet were summarised. A frequency distribution of landslide ponds by size class was calculated and graphed. The total number of landslides for each landslide type and the total number of pond-bearing landslides for each landslide type were then calculated and compared in a paired graph. Mean pond size overall, mean pond size per landslide type, and mean pond size per geomorphic location within landslides were then calculated and graphed. Minimum, maximum, and median pond sizes were also determined for each of the preceding categories. Due to time constraints, no attempt was made to map and analyse ponds on undisturbed terrain. 4.4. Results 4.4.1 Pond characteristics in overall study area Of the total 1,638 landslides identified in the study area, 223 landslides with ponds were identified, and 755 ponds were recorded on these pond-bearing landslides (Table 4-1). There appeared to be a clustering pattern, with some large landslides containing many small ponds. Other smaller landslides only had a few ponds. Many of the ponds appeared to be persistent, 177 with signs of cattails (Typha spp. – most likely Typha latifolia) and development into wetland ecosystems. Ponds occupied only a small fraction of the total landslide area. The total area of landslides in the study area was 768.25 km2 (76,825 ha), including some added area not digitised in Severin’s work. The total area of ponds on these landslides was 111.647 ha. Therefore, landslide ponds only occupied 0.14% of the total landslide area. The minimum landslide pond size was 0.0009 ha while the maximum pond size was 5.890 ha, showing a wide range of sizes. However, most ponds were in the <1.00 ha size class (Figure 4-4). The mean pond size was 0.15 ha, and the median pond size was 0.05 ha. Because the landslides in Severin’s (2004) work were not digitised as enclosed polygons with a fixed area, it was not possible to calculate the area of pond-bearing landslides separately for the present study. Therefore, calculation of the proportion of total area occupied by ponds on just pond-bearing landslides was not possible. Re-digitising 1,600 landslides would have added significantly to the workload for this study and was not feasible with the time constraints and scope of the project. 178 Table 4-1. Summary information on ponds and landslides in the 768.25 km2 (76,825 ha) study area. Total number of landslides mapped 1,638 Total number of landslides with ponds 223 Percent (%) of total landslides that contain ponds 13.61 Total number of ponds on landslides 755 Mean number of ponds/pond-bearing landslide 3.386 Minimum number of ponds/pond-bearing 1 landslide Maximum number of ponds/pond-bearing 29 landslide Total pond area (ha) on landslides 111.647 Minimum landslide pond size (ha) 0.0009 Maximum landslide pond size (ha) 5.890 Mean landslide pond size (ha) 0.148 Total landslide area (ha) 76,918.527 Percentage of total landslide area covered by 0.14 ponds 179 Figure 4-4. Frequency distribution graphs showing landslide pond size distribution in hectares (ha). The upper graph shows all landslide-generated ponds in Mapsheet 94A, while the lower graph shows the size distribution for landslide ponds ≤ 1.00 ha. 4.4.1.1 Pond number and total area by geomorphic location Over the study area, the majority of the 755 ponds were located on the body of landslides (370 ponds, 49%), followed by the toe (236 ponds, 31%) and then the head (149 ponds, 20%). (Figure 4-5). 180 Figure 4-5. Total number of landslide ponds per geomorphic location. Ponds on the body of the slides also comprised the highest total area (53.252 ha), which represented almost 48% of the total pond area (Figure 4-6). This was followed by pond area on the toe (27%) and pond area on the head (25%). Figure 4-6. Total landslide pond area (ha) by geomorphic location. 181 4.4.1.2 Pond size by geomorphic location Pond size was quite variable both within and among different geomorphic location types. Ponds on the head had the largest average size at 0.186 ha, while ponds on the toe were the smallest average size at 0.130 ha (Figure 4-7). The average size of ponds on the body was 0.144 ha. Overall, a location on the body contained a pond with the smallest size (0.0009 ha) and a location on the body also contained the largest pond size (5.890 ha). The head had the largest range of pond sizes (1.569 ha), while the body had the smallest range (0.059 ha). The range of pond sizes on the toe was (0.356 ha). Figure 4-7. Mean pond size (ha) by geomorphic location on landslide. 4.4.1.3 Landslide types for slides containing ponds Landslide ponds occurred more frequently on certain landslide types (Figure 4-8). The highest number of pond-bearing slides were retrogressive rotational failures, followed by rotational failures and then compound failures. Proportionally, retrogressive rotational 182 failures comprised approximately 39.5% of the total landslides with ponds, followed by rotational failures at 32.7%, and then compound failures (14.8%). Figure 4-8. Paired graph showing overall number of landslides per landslide type compared to number of pondbearing landslides per landslide type. Landslide type names have been abbreviated to accommodate graph. Type abbreviations: CO = Compound; EF = Earth flow; MF = Mobile flow; MR = Multi-level rotational; RA = Ravelling; RE = Retrogressive rotational; RO = Rotational; SR = Shallow retrogressive. Note: No ponds were found on Ravelling (RA) or Earth flow (EF) landslide types. 4.4.2 Ponds and landslide type 4.4.2.1 Number of ponds per landslide type Pond numbers ranged widely across landslide types (Figure 4-9). Retrogressive rotational failures contained by far the greatest overall number and proportion of the total 755 ponds (394 ponds, approximately 52%). The next highest proportion of ponds was on rotational failures (20%, 151 ponds), followed by compound failures (14%, 107 ponds). Shallow retrogressive failures had the lowest proportion of total ponds (approximately 2%, 13 ponds). 183 Figure 4-9. Number of ponds per landslide type. 4.4.2.2 Pond area per landslide type Within the study area, the planimetric surface area occupied by landslide ponds primarily occurred in one landslide type (Figure 4-10). By far, retrogressive rotational failures contained the highest total area (77.696 ha, 69.59 %), followed distantly by rotational failures (18.877 ha, 16.91 %) and then compound failures (5.737 ha, 5.14 %). Figure 4-10. Pond area (ha) in each pond-bearing landslide type. 184 4.4.2.3 Mean pond size per landslide type Pond size varied widely between landslide types in the study area (Figure 4-11 and Appendix 8). Retrogressive rotational failures had the largest average pond size (0.197 ha), followed by rotational failures (0.125 ha), and then multi-level rotational failures (0.102 ha). Shallow retrogressive slides had the smallest average pond size (0.051 ha). There was quite a range of pond sizes both among and within slide types, with large standard deviations. Retrogressive rotational failures had the smallest minimum pond size (0.0009 ha), as well as the largest maximum pond size (5.890 ha). Mobile flows had the largest minimum pond size (0.005 ha). Shallow retrogressive failures had the smallest maximum pond size (0.287 ha). Within slide types, retrogressive rotational failures had the greatest pond size range (5.889 ha), followed by rotational failures (1.651 ha). Shallow retrogressive rotational failures had the smallest pond size range (0.285 ha). However, it should be noted that there were only a few shallow retrogressive rotational failures in the data set. The next smallest pond size range was within compound failures, at 0.290 ha. Figure 4-11. Mean pond size (ha) per landslide type. 185 4.4.3 Geomorphic location of ponds and landslide type This section presents the results of compilation and analysis of pond area and size for various combinations of pond geomorphic locations and landslide types. In all cases for the results regarding pond size, the standard deviations were quite large, often much larger than the average size (see Appendix 9). 4.4.3.1 Head and landslide type 4.4.3.1a Number and proportion of ponds on head per landslide type Ponds on the head of landslides occurred much more frequently on some landslide types than on others (Figure 4-12). The highest number and proportion of ponds on the head were on retrogressive rotational failures (80 ponds, 54%). Rotational failures had the next highest number and proportion of ponds (29 ponds, 19%), followed by multi-level rotational failures. The smallest number and proportion of ponds on the head occurred on the single mobile flow on which landslide ponds were detected. Figure 4-12. Number of ponds on head per landslide type. 186 4.4.3.1b Pond area on head per landslide type The area occupied by ponds on the head of slides varied strongly by landslide type (Figure 413). Most of the total area of ponds on the head of slides was represented by retrogressive rotational failures, comprising 22.493 ha (81.21 %). The second highest area was on rotational failures, with a much smaller value of 2.377 ha (8.58 %), followed by compound failures (1.302 ha, 4.70 %). Figure 4-13. Pond area (ha) on head per landslide type. 4.4.3.2 Body and landslide type 4.4.3.2a Number and proportion of ponds on body per landslide type Occurrence of ponds on the body of landslides varied but showed a tendency towards certain landslide types (Figure 4-14). With a similar trend as for ponds on the head, the highest number and proportion of ponds on the body occurred on retrogressive rotational failures (192 ponds, 50.26%). Rotational failures had the second highest number and proportion of 187 ponds (81 ponds, 21.2 %). Shallow retrogressive failures represented the smallest number and proportion of ponds on the body (5 ponds, 1.31%). Figure 4-14. Number of ponds on body per landslide type. 4.4.3.2b Pond area on body per slide type The area occupied by ponds on the body of slides showed a strong tendency toward just one or two landslide types in the study area (Figure 4-15). Retrogressive rotational failures contained the highest overall area of ponds on the body, at 36.946 ha (69.38 %). Rotational failures were a distant second at 9.983 ha (18.75 %), followed by compound failures (2.849 ha, 5.35 %). 188 Figure 4-15. Total pond area (ha) on body per landslide type. 4.4.3.3 Toe and landslide type 4.4.3.3a Number and proportion of ponds on toe per landslide type The occurrence of ponds on the toe of a landslide was variable, but more evenly distributed among landslide types than that of ponds on the body (Figure 4-16). More than half of ponds on the slide toe occurred on retrogressive rotational failures (122 ponds, 51.69%). The next highest number of ponds on the toe was represented by rotational failures (41 ponds,17.37%), similar to the trend for ponds on the head and body. No ponds on the toe were located on mobile flows, and the next lowest pond number on the toe was found on shallow retrogressive failures (3 ponds, 1.27%). 189 Figure 4-16. Number of ponds on toe per landslide type. 4.4.3.3b Pond area on toe per landslide type Total area of ponds on the toe of landslides was somewhat variable between landslide types (Figure 4-17). The pond area on the toe was highest on retrogressive rotational failures, at 18.257 ha (59.47 %), and this slide type also had the greatest overall pond area in the study (see Figures 4-13 and 4-14). The second highest area of ponds on the toe was on rotational failures (6.517 ha, 21.23 %), followed by multi-level rotational failures (4.124 ha, 13.43 %). 190 Figure 4-17. Pond area (ha) on toe per landslide type. 4.4.4 Landslide pond development The landslide-generated ponds in the study area exhibited varying stages of development, both between and within individual landslides. Because of the seasonal nature of pond water levels and vegetation, it would not have been accurate to attempt to categorise the ponds in the study according to development stage. However, some representative photographs illustrate the diversity of vegetation development present on landslide-bearing ponds in the mapsheet study area. These photographs were taken during fieldwork for Chapters 2 and 3 in the same area. Figure 4-18 shows a pond recently formed in fresh unvegetated soil. Figure 4-19 shows a pond with a more stable bank, with vegetation growing around the perimeter of the pond. Figure 4-20 shows a pond with an obviously stable bank, surrounded by vegetation, and containing aquatic vegetation including cattail (Typha spp. – most likely Typha latifolia). 191 Figure 4-18. Very new pond on landslide (Beatton River Landslide). The most recent major movement on the landslide occurred in 2015, so the pond was approximately three years old at time of photograph. (Photo June 19, 2018) Figure 4-19. Newer pond on landslide (Beatton River Landslide). Pond is on an older part of the landslide near the toe. (Photo August 9, 2017) 192 Figure 4-20. Persistent pond on landslide (Cecil Lake Landslide). Landslide occurred in 1998, thus the pond was approximately 20 years old at time of photograph. (Photo June 23, 2017) 4.5. Discussion This study set out to compile an inventory of landslide ponds on NTS Mapsheet 94A and describe area, size, and distribution of the ponds in relation to overall number and area of landslides, geomorphic locations on the landslide, and occurrence on landslide types. The overarching purpose was to employ this information to identify potential ecological implications of pond occurrence and distribution and provide recommendations for management. 193 4.5.1 Landslide pond characteristics and distribution The results showed a wide range of pond sizes, with the average size of ponds for each landslide type falling within an intermediate range. Research has shown that intermediate sized ponds (ranging in size from 200 to 4000 m2, or 0.02 to 0.4 ha) contain the highest density, richness, and diversity of pond-breeding amphibians (Semlitsch et al. 2015). In the present study, 497 ponds (over 65%) fell within the intermediate size range. This could be significant for maintenance of amphibian diversity in the study area. Although no amphibians or aquatic insects were observed in the ponds encountered on the Cecil Lake, Beatton River, and Hasler Flats landslides (Chapters 2 and 3), waterfowl were seen on some of the ponds. The ponds in the study area also exhibited a range of stages of evolution. This diversity of developmental stages could allow for a variety of different macroinvertebrate communities to develop, each taking advantage of the particular hydrologic, sedimentary, and vegetative conditions present (Jeffries 2011). The characteristics of each geomorphic location on the landslide may influence the distribution of ponds. Generally, each part of a landslide has different kinds and orientations of geomorphic structures (Parise 2003). In the present study, ponds were most prevalent on the body of landslides, followed by the toe. Ponds on the body represented the highest total number, as well as the highest total area. Greater pond presence on the landslide body may be due to the diversity in topography that results as the slide material is moved, creating depressions and cutting off drainage. As the landslide stabilises, vegetation starts to encroach, further stabilising ponds. The conditions are generally better for plants to persist on the body compared to the steeper nature of the head, which prevents soil from building up and hinders plant root establishment and persistence (Walker and Shiels 2013). 194 As discussed in the Introduction, a network of varying sizes of ponds can provide value for dispersal of water-dependent organisms (Pop and Chitu 2013). Ponds on the toe represented the second highest number and total area of all the mapped ponds but had the smallest average size. These toe ponds had a size range that was less than the head and greater than the body, and thus had greater size diversity than ponds on the body. This size variation could be due to the broken up and uneven nature of the terrain often present on the toe, where large amounts of material are deposited, often rapidly. Although ponds were more prevalent on the landslide body, the head had the largest maximum pond size and the greatest size range. Most ponds on the head of the landslide were at or near the transition zone with the body. Deep, long depressions were more likely to be present at this sharp transition in slope from positive to negative, forming the larger ponds. The smaller ponds on the head tended to be in small depressions. These findings suggest that the head of a landslide may provide more diverse habitats for some aquatic organisms, as well as for larger animals seeking water or shade. The apparent relationship between landslide type and number of ponds in the study area may be a function of the underlying material and the topographical characteristics of the landslide. Most of the ponds in the data set were located on either retrogressive rotational slides or rotational slides, followed by compound slides. By far, most ponds were found on retrogressive rotational slides. The persistence of ponds on this landslide type may be due to pre-existing fault planes that form depressions and restrict drainage following movement of slide material. Retrogressive rotational slides have multiple weak layers and fractures (Severin 2004), creating many potential sites for ponds. Ponds on retrogressive rotational slides also had the greatest mean size and the greatest range of sizes, followed by ponds on 195 rotational slides. Retrogressive rotational slides tend to produce a series of geomorphic features as the land mass slumps and shifts. This process may result in a great variation of depressions and cracks where ponds can form. Rotational failures have a similar configuration, but with fewer fracture planes (Cruden and Varnes 1996; Severin 2004). The compound failures possibly have a relatively high proportion of ponds in the study because the overall slope on this type of landslide is gentler, enhancing the ability of the material to retain water. The fewest number of ponds were on shallow retrogressive failures. It is possible the fracture planes of this landslide type were not severe enough to create persistent depressions for water retention. Geomorphic location of ponds on the landslide varied somewhat with landslide type. For ponds on the head, by far the highest number and total area were on retrogressive rotational failures, followed by rotational failures and then compound failures. This prevalence on retrogressive rotational failures could be due to the relatively steep headscarp of these types of landslides and the associated deep and wide fissures at the transition zone with the body. Regarding ponds on the body, the highest number and area of ponds was again on retrogressive rotational failures, followed by rotational failures and then shallow retrogressive failures. Total pond area on the body was higher than for ponds on the head for most landslide types noted. For ponds on the toe, the trend was the same, with the highest number and area of toe ponds occurring on retrogressive rotational failures and rotational failures. Retrogressive rotational failures have multiple shear zones (Cruden and Varnes 1996), which can produce depressions and fissures at all geomorphic locations of the landslide. Overall, the results suggest that retrogressive rotational failures are more 196 ecologically diverse than the other landslides sampled regarding pond size, area, distribution, and persistence. 4.5.2 Landslide pond dynamics Although ponds on landslides can persist for many decades, they evolve and change over this time. The ponds in the study area showed a diversity of stages of evolution. Initially, when most ponds form after a landslide event, they are in fresh unvegetated soil (see Figure 4-18). Over time, the vegetation starts to grow in around the pond from propagules either within the soil or from surrounding rafts of vegetation or from seeds dispersed from adjacent forest vegetation (see Figure 4-19). This vegetation can serve to stabilise the pond, and it also can influence the ecology of the site. Eventually, the area surrounding the pond may become fully vegetated and the pond may persist for years (see Figure 4-20). As habitat and water persistence change, so do the populations and compositions of plants and animals. For example, macroinvertebrate communities develop and change in response to pond persistence and hydrological cycles, and at times there is a fine threshold between different community compositions (Jeffries 2011). The gradual revegetation of the pond site can also influence the size and persistence of the pond. Surrounding vegetation as well as aquatic vegetation can shrink the pond either seasonally or over years, as established vegetation takes up the water through transpiration, eventually lowering the overall reserves. 4.5.3 Beaver influence on landslide ponds Evidence on the study sites visited for field sampling in Chapters 2 and 3 suggested that beavers can play a part in forming or altering ponds on landslides. The North American beaver (Castor canadensis) is common in the study area and throughout northeastern British 197 Columbia and can significantly impact landscapes that have water present. Beavers can alter hydrogeomorphic and ecological processes through dam building and the associated felling of trees, and excavation and transport of large amounts of sediments, resulting in the flooding of various terrains (Butler and Malanson 1995; Westbrook et al. 2017). Beaver-modified landscape patches produce distinct habitats that can increase richness and abundance of terrestrial and semi-aquatic mammals (Nummi et al. 2019). Although it was difficult to identify beaver ponds in the imagery used, in the field there was evidence of beaver activity on the Cecil Lake landslide study site (Figures 4-21, 4-22, and 4-23), as well as the Hasler Flats landslide study site (Figure 4-24) sampled in work for Chapters 2 and 3. The beavers appear to have significantly influenced the configuration and size of the bigger ponds on site at the Cecil Lake landslide (Figure 4-22). Field evidence of old weathered, advanced-decay gnawed logs and stumps (Figure 4-23) indicate beavers have maintained a presence for many years and operated in cycles on different parts of the landslide. In many areas on the Cecil Lake landslide there were small beaver-browsed sapling stumps and beaver trails throughout the woods leading to ponds. On the Hasler Flats landslide, there was evidence of very recent felling of large aspen trees in addition to well-worn trails leading from the sidescarp to the landslide ponds (Figure 4-24). 198 Figure 4-21. Recent beaver gnawing activity on young sapling near pond (Cecil Lake Landslide). (Photo September 7, 2017) Figure 4-22. Beaver pond on the Cecil Lake Landslide. (Photo June 22, 2017) 199 Figure 4-23. Older beaver gnawing activity on the Cecil Lake Landslide. (Photo September 10, 2017) 200 Figure 4-24. Recent beaver activity at the Hasler Flats landslide. Top image shows well-used beaver path (bottom centre of photo) leading from the sidescarp to a landslide pond. Lower image shows very recent felling of large aspen trees (Populus tremuloides) by beavers just above the sidescarp. (Photos August 14, 2018) 201 4.5.4 Limitations of the study A limitation to the landslide ponds study was the intermediate and variable resolution of the imagery used when determining and mapping landslide ponds. Some ponds may have been missed if they were too small, or if the available imagery in the area was of a poorer resolution. However, the vast majority of ponds present were most likely identified and digitised, and any missed ponds probably did not affect the overall results. Another limitation of the study was the fact that the pre-existing digitised landslides did not have individual areas assigned to them in a format that could be georeferenced. Thus, calculation of proportions of total area of pond-bearing landslides occupied by ponds was not possible. This information would have been a useful metric to have for comparison and consideration. However, the landslide area summary information that was available provided an initial baseline for calculating the proportion of total landslide area occupied by ponds. The georeferenced and numbered landslide type designations that Severin (2004) assigned to each landslide were also unavailable, so in this study I assigned my own classifications for each individual pond-bearing landslide based on Severin’s definitions. Landslide types were classified mainly using Severin’s geomorphic symbols and the DEM imagery. My designations may not have been completely accurate, as I did not have access to high resolution aerial photographs as stereo pairs and am not a trained geomorphologist. Nevertheless, the classifications were likely accurate enough for distinguishing between obviously different landslide types and describing pond distribution. 202 An additional limitation to the study was the lack of previous research on landslide ponds, for comparison. There have not been many studies to inventory landslide ponds, let alone to describe ecological elements. This study is thus a baseline for future research. 4.5. Conclusions and recommendations In this study I provided an initial inventory of ponds on landslides in an area of northeastern British Columbia that is particularly susceptible to landslides. I also presented details on the number, size, and distribution of ponds on different geomorphic locations within landslides and among different landslide types. Although they only occupy a small proportion of the total landslide area in the mapsheet study area, landslide ponds may provide important ecological roles for amphibians, ungulates, birds, and other wildlife both at the local and landscape scales in the Peace River Region. These ponds serve many purposes, including nesting, feeding, shelter, water, protection, connectivity, and biodiversity. Pond size and distribution impact wildlife species richness and overall diversity, as an array of smaller ponds yields a greater number of species and higher conservation value than a large pond of the same total area (Oertli et al. 2002). This study provides the first detailed baseline information on landslide pond distribution in the Peace River Region and adds a valuable component to the knowledge base on water bodies in the area. The findings present a benchmark for conservation considerations in the Region. As landslide ponds do not occur in great numbers on the landscape here and yet are potentially high in ecological value, efforts should be made to conserve them. Conservation would allow for preservation of breeding, nesting, feeding, and shelter habitats for various species, as well as connectivity for migrating species. Knowledge about the specific 203 ecological value of these landslide ponds is significantly lacking and requires further investigation. To better understand how these ponds are used by wildlife, a subset of the pond-bearing landslides should be selected to sample in the field. Ecological information such as pond plant species composition, aquatic invertebrates, and signs of wildlife use should be recorded in detail for each pond in the subset. Ideally, such a study would be carried out over multiple seasons and years to capture the full breadth of use of the pond, as well as any changes to the pond. It would also be informative to carry out similar research in landscapes dominated by other landslide types such as rock falls. Although the ecological value of landslide ponds is recognised, geohazard assessment considers the presence of pooled water on unstable slopes a dangerous situation to be avoided. In fact, some management measures recommend draining ponds on landslides (Kansas Geological Survey 1999). Therefore, a balance must be sought between conserving important habitat and preventing catastrophic reactivation of landslides. Landslide ponds may serve an important role as an indicator for land management decisions concerning infrastructure, resource development, home building, and other activities on the land base. Persistent ponds signify a high water table and soil saturation, conditions which can indicate the potential for slope instability and increased possibility of flooding. Care should be taken to develop away from areas where landslide ponds are present. If landslides with ponds already exist near developed areas or sensitive fisheries habitat, they should be monitored on a regular basis and assessed for reactivation. It is possible some ponds should be drained if there are signs of imminent danger of slope movement. For landslides in remote areas, however, the ponds should be left intact to provide connectivity, habitat, and other ecological roles. 204 4.6 References ASTER GDEM v2. 2021. Advanced Spaceborne Thermal Emission and Reflection Radiometer (ASTER) Global Digital Elevation Map. https://asterweb.jpl.nasa.gov/gdem.asp Bobrowsky, P.T., N. Catto, and V. Levson. 1990. Reconnaissance Quaternary geological investigations in Peace River District, British Columbia (93P, 94A). Geological Fieldwork, Paper 1991-1. Bobrowsky, P.T., and N.W. Rutter. 1992. The Quaternary geologic history of the Canadian Rocky Mountains. Geographie physique et Quaternaire, 46(1): 5-50. Butler, D.R., and G.P. Malanson. 1995. Sedimentation rates and patterns in beaver ponds in a mountain environment. Geomorphology, 13: 255-269. Catto, N.R. 1991. Quaternary geology and landforms of the eastern Peace River region, British Columbia NTS 94A/1, 2, 7, 8. Province of British Columbia, Ministry of Energy, Mines and Petroleum Resources, Mineral Resources Division, Geological Survey Branch, Open File 1991-11, 23 pages. Catto, N., D.G.E. Liverman, P.T. Bobrowsky, and N.W. Rutter. 1996. Laurentide, Cordilleran, and Montane glaciation in the Western Peace River – Grande Prairie Region, Alberta and British Columbia, Canada. Quaternary International, 32: 21-32. Coe, J.A., W.L. Ellis, J.W. Godt, W.Z. Savage, J.E. Savage, J.A. Michael, J.D. Kibler, P.S. Powers, D.J. Lidke, and S. Debray. 2003. Seasonal movement of the Slumgullion landslide determined from Global Positioning System surveys and field instrumentation, July 1998 – March 2002. Engineering Geology, 68: 67-101. Coe, J.A., J.P. McKenna, J.W. Godt, and R.L. Baum. 2009. Basal-topographic control of stationary ponds on a continuously moving landslide. Earth Surface Processes and Landforms, 34: 264-279. Cruden, D.M., Z.-Y. Lu, and S. Thomson. 1997. The 1939 Montagneuse River landslide, Alberta. Canadian Geotechnical Journal, 34: 799-810. Cruden, D., and D. Varnes. 1996. Landslide types and processes. In: Turner, A., Schuster, R. (eds.), Special Report 247: Landslides Investigation and Mitigation. National Research Council, Transportation Research Board, Washington, DC, pp. 36-75. Dandurand, R. 2018. Investigating regional groundwater flow influences on slope stability in unlithified materials. Unpublished Master of Science Thesis, Simon Fraser University, 139 pages. Fletcher, L., O. Hungr, and S.G. Evans. 2002. Contrasting failure behaviour of two large landslides in clay and silt. Canadian Geotechnical Journal, 39: 46-62. Geertsema, M., and J. Pojar. 2007. Influence of landslides on biophysical diversity - A perspective from British Columbia, Geomorphology, 89: 55-69. Global Mapper. 2021. https://www.bluemarblegeo.com/global-mapper/ 205 Google Earth Pro. 2021. https://www.google.com/earth/versions/ Hartman, G.M.D. 2005. Quaternary stratigraphy and geologic history of the Charlie Lake (NTS 94A) map-area, British Columbia. M.Sc. thesis, Simon Fraser University, Burnaby, B.C., 165 pp. Hartman, G.M.D., and J.J. Clague. 2008. Quaternary stratigraphy and glacial history of the Peace River valley, northeast British Columbia. Canadian Journal of Earth Science, 45: 549564. Highland, L.M., and P. Bobrowsky. 2008. The landslide handbook – A guide to understanding landslides. Reston, Virginia, U.S. Geological Survey Circular 1325. 129 pp. Hutchinson, J.N. 1988. General Report: Morphological and geotechnical parameters of landslides in relation to geology and hydrogeology. In Proceedings, 5th International Symposium on Landslides, Lausanne, Vol. 1, pp. 3-35. Jeffries, M.J. 2011. The temporal dynamics of temporary pond macroinvertebrate communities over a 10-year period. Hydrobiologia, 661:391-405. Kansas Geological Survey. 1999. Public Information Circular (PIC) 13. Kansas Geological Survey, Public Outreach. http://www.kgs.ku.edu/Publications/pic13/pic13_5.html Krueger, K.A., and B.G. Johnson. 2016. The effect of sub-alpine landslides on headwater stream gradient and beaver habitat. Physical Geography, 37(5): 344-360. DOI: 10.1080/02723646.2016.1218723 Mathews, W.H. 1978. Quaternary stratigraphy and geomorphology of Charlie Lake (94A) map-area, British Columbia. Geological Survey of Canada, Paper 76-20, 25 pages. Mathews, W.H. 1980. Retreat of the last ice sheets in northeastern British Columbia and adjacent Alberta. Geological Survey of Canada Bulletin 331, 22 pages. Meidinger, D., and J. Pojar. (Eds.) 1991. Ecosystems of British Columbia (342 p). Special Report Series 6, Victoria, BC: B.C. Ministry of Forests and Range Research Branch. National Topographic System (NTS). 2021. Website. https://naturalresources.canada.ca/earth-sciences/geography/topographic-information/maps/nationaltopographic-system-maps/9767 Nummi, P., W. Liao, O. Huet, and E. Scarpulla. 2019. The beaver facilitates species richness and abundance of terrestrial and semi-aquatic mammals. Global Ecology and Conservation, 20: 1-10. Parise, M. 2003. Observation of surface features on an active landslide, and implications for understanding its history of movement. Natural Hazards and Earth System Sciences, 3: 569580. Pop, D.E., and Z. Chitu. 2013. Landslides and biodiversity conservation: the importance of an integrated approach. A case study: the Subcarpathian part of the Doftana watershed (Prahova County, Romania). Revista de geomorfologie, 2: 57-68. 206 Sasaki, N., and T. Sugai. 2015. Distribution and development processes of wetlands on landslides in the Hachimantai volcanic group, NE Japan. Geographical Review of Japan Series 8, 87(2):103-114. Semlitsch, R.D., W.E. Peterman, T.L. Anderson, D.L. Drake, and B.H. Ousterhout. 2015. Intermediate pond sizes contain the highest density, richness, and diversity of pond-breeding amphibians. PLoS ONE 10(4): e0123055. doi:10.1371/journal.pone.0123055 Severin, J.M. 2004. Landslides in the Charlie Lake map sheet, Fort St. John. Unpublished Master’s thesis, University of British Columbia, 395 pp. Shapley, M.D., B.P. Finney, and C.R. Krueger. 2019. Characteristics of landslide-formed lakes of central Idaho: high-resolution archives of watershed productivity and clastic sediment delivery. Geological Society of America, Special Paper 536, pp 241-258. Varnes, D.J. 1978. Slope movement types and processes. In Landslides – Analysis and Control. Special Report of the National Research council and Transportation Research Board, 176. Edited by R.L. Schuster and R.J. Krizek. pp. 11-33. Van Esch, K.J.B. 2012. Failure behaviour of bedrock and overburden landslides of the Peace River Valley near Fort St. John, British Columbia (T). University of British Columbia. Retrieved from https://open.library.ubc.ca/collections/ubctheses/24/items/1.0072987 Walker, L.R., and A.B. Shiels. 2013. Landslide Ecology. Cambridge University Press, New York. 300 pages. Westbrook, C.J., D.J. Cooper, and C.B. Anderson. 2017. Alteration of hydrogeomorphic processes by invasive beavers in southern South America. Science of the Total Environment, 574: 183-190. World Imagery. 2021. https://www.arcgis.com/home/item.html?id=10df2279f9684e4a9f6a7f08febac2a 207 Chapter 5. Conclusions and Recommendations for Landslide Recovery and Management 5.1 Conclusions This research set out to describe and quantify various aspects of biophysical diversity on landslides in the landslide-prone Peace River Region of northeastern British Columbia and present ecological implications and management recommendations based on the findings. The central research questions for this research were: (1) Are landslides demonstrably more biophysically diverse than undisturbed ecosystems? (2) To what extent do landslides rearrange the relative abundance of site series/types on a slope compared to adjacent undisturbed terrain? (3) What is the extent turnover in microsite and plant species diversity on landslides, and how does this compare to adjacent undisturbed terrain? (4) Is vegetation diversity on landslides significantly related to geomorphological diversity? (5) What is the distribution and abundance of landslide ponds at a regional and local scale, and what are the ecological and management implications? The overarching purpose of the work was to compare biophysical diversity within and among landslides, and between landslides and surrounding undisturbed terrain. An additional objective was to investigate possible correlations between vegetation diversity and site diversity. Vegetation and site characteristics on three different landslides were measured and analysed for alpha diversity and beta diversity values (Chapter 2 and Chapter 3, respectively), and characteristics of ponds on landslides within a larger area in the region were also assessed, described, and analysed (Chapter 4). The findings of this study revealed that 208 quantification and prediction of biophysical diversity on a severe disturbance such as a landslide is complex and highly dependent on individual study site characteristics. Despite these challenges, it is still possible to analyse the results and draw learnings for application in management, ecology, and rehabilitation on landslides in the Peace River Region and for landslides at the broader scale. This work showed that following a mass movement in the Peace River Region, plant community composition on landslides varies depending on the age and size of the landslide and the slope and soil development of the various geomorphological features present. Exotic forb species tend to dominate in early stages of landslide revegetation and can persist if the disturbance periodically reactivates, effectively preventing the establishment of shrubs and trees. The landslides in the study were overall less diverse in alpha diversity of plant communities than the surrounding undisturbed terrain, a finding which diverged from initial expectations. However, the landslides were more diverse in abundance and distribution of site types/series and geomorphic features than the undisturbed terrain. Although there was a greater proportion of mesic sites on landslides, there were also more extreme site series on a scale of moisture regime. Geomorphic types were overall more diverse on the landslides due to mass movement and substrate rearrangement, and type diversity and surface roughness both tended to decrease with age of the landslide. Therefore, although the landslides were lower in plant alpha diversity, the site series and geomorphic diversity present provide conditions for a greater variety of plant communities and wildlife habitats over time. 209 Beta diversity often reflects the health and proper functioning of an ecosystem. Beta diversity or turnover of both vegetation and environment variables was generally higher on landslides than the surrounding undisturbed terrain, with a strong relationship between the vegetation and environment beta diversities. Beta diversity was also affected by spatial and temporal characteristics of the study area. The largest and oldest landslide, Cecil Lake, showed the highest mean vegetation beta diversity and also the most variable vegetation beta diversity. This suggests that processes affecting turnover of plant communities may be at work to varying degrees on different parts of a landslide within a given time period. The vegetation beta diversity on the surrounding terrain was both lower and much less variable, suggesting a state of relative stability. The smallest landslide, Hasler Flats, had the highest beta diversity but also had much less variability. This suggests size can inversely influence beta diversity, with smaller areas facilitating greater relative patchiness. Overall, environment beta diversity was much lower than vegetation beta diversity for all study sites, and it was also lower on undisturbed areas compared to the landslides. Cecil Lake and Hasler Flats landslides both had environment beta diversity values that were nearly twice as much as the youngest landslide, Beatton River, but Cecil Lake had more than twice the variability as Hasler Flats. Beatton River had a markedly higher variability. Interestingly, Beatton River had some higher vegetation and environment alpha diversity values than the oldest landslide, Cecil Lake. The above findings highlight the variability of the effects of patchiness on biophysical diversity in general. This study of landslides in the Peace River Region uncovered variable relationships between vegetation and site or environment diversity, depending on the spatial and temporal scale of the samples. The NMS ordination analysis on the BEC vegetation plots showed some weak 210 relationships between plant community development and slope gradient, and to a lesser degree mesoslope position, heat load index, and moisture. Contrary to expectations, a significant relationship was not found between vegetation alpha diversity and environment (geomorphic) alpha diversity on any of the three landslides studied (Chapter 2). However, there was a significant positive relationship between vegetation beta diversity and environment beta diversity on the transects sampled in Chapter 3. These findings suggest that while within-plot vegetation diversity seems to be independent of within-plot environmental diversity, vegetation turnover over space is distinctly linked to microsite condition turnover. The complex nature of these relationships between vegetation and environment indicates a scale-dependency that is not yet clear and requires further investigation. In general, it appears that during the first 20 or more years following landslide occurrence, plant community succession is still sorting out, at the same time as the terrain is weathering and evolving. Although both prehistoric and historic landslides are abundant along the Peace River and its tributaries, the research in Chapter 4 found only a small proportion of these landslides contained persistent ponds. The results showed trends in landslide pond size, geomorphic location on the landslide, and association with certain landslide types. Most ponds were under one hectare in surface area, with most being less than one-tenth that size. Further, these ponds tended to occur more frequently and in greater numbers on certain types of landslides, namely retrogressive rotational, rotational, and multi-level rotational landslides. In addition, ponds occurred in greater concentrations on the body of the landslide, followed by the toe and then least frequently on the head. However, ponds on average tended to be larger on the head of landslides. This is likely due to the presence of the rupture zone below 211 the headscarp, where movement of material away from the zone followed by stabilisation can cause large depressions, restricting drainage. Landslide ponds encountered during fieldwork for Chapter 2 and Chapter 3 had evidence of beaver activity and alteration, as well as use by waterfowl and other animals. 5.2 Recommendations The findings of this research provide a foundation to begin managing landslides in the Peace River Region from an ecological perspective, considering succession and the influence of spatial and temporal scales. Landslides close to communities, infrastructure, or other important developments or ecosystems should be the priority for management and restoration. Landslides abutting large rivers used for drinking water or important fishbearing streams should be given special attention, as sediment input can be substantial. The type of landslide and its geomorphological characteristics may also provide guidance for prioritising management. Rotational landslides tend to be more susceptible to reactivation, especially those with steeper slopes of perhaps 20 degrees or more. The presence of moving water within the landslide also tends to reactivate portions of the slide. The Beatton River landslide had some areas of seepage and debris flow, while the Cecil Lake landslide was influenced by a creek running through it from the south, creating an active gully. When landslides first occur, they should be assessed in the field to determine potential risks for reactivation. Once a landslide has been assessed and is determined safe to work on, initial measures should focus on revegetating and stabilising the terrain to enhance the rate of ecological succession. Invasive, exotic plant species such as Melilotus spp. and Sonchus spp. are obviously very effective at colonising, stabilising, and enriching freshly disturbed terrain. 212 However, these species pose the problem of creating reduced plant community diversity and structure over time. To alleviate this, land managers should also plant a variety of competitive, rapidly growing native pioneer grass seed mixes such as blue wildrye (Elymus glaucus) and Canada bluejoint (Calamagrostis canadensis), forbs including wild sarsaparilla (Aralia nudicaulis), palmate coltsfoot (Petasites frigidus), showy aster (Aster conspicuus) and false Solomon’s seal (Smilacina racemosa), and common shrubs prickly rose (Rosa acicularis), saskatoon (Amelanchier alnifolia), alder species (Alnus spp.), red-osier dogwood (Cornus stolonifera), and highbush cranberry (Viburnum edule), as found most abundantly on the landslides in this study. In addition, ecologically suitable deep-rooted or high evapotranspiration trees such as trembling aspen (Populus tremuloides), white spruce (Picea glauca) and cottonwood (Populus balsamifera) should be planted where possible and safe to do so to help stabilise the soil and take up extra moisture. Decisions on species selection should incorporate local and regional goals for landscape ecosystem health and measures around climate change adaptation. Organic amendments and bird perches in open areas may also be necessary to ensure adequate substrate conditions, plant dispersal and establishment. Further, these sites should be monitored over time, and managers should be prepared to replant or stabilise the slope if steep portions of the landslide are reactivated. It is also important to note that there are often other disturbances interacting with landslides, such as floods and fires. These disturbances should also be taken into consideration when planning and monitoring restoration. Because of the complexities of landslide succession and the potential for reactivation, restoration on landslides should focus on ecosystem recovery and biodiversity, rather than species composition. Ecosystem recovery is not intended to return the disturbed system to its 213 historic condition, but rather to its historic trajectory. Work should involve stabilising the soil through planting and bioengineering measures, rehabilitating water courses, and creating a diverse multi-level vegetation cover of mostly native species. The measures described above could be applicable to deep-seated, moderate to steep landslides occurring on glaciated sites in unconsolidated glaciolacustrine material elsewhere in British Columbia and in other parts of the world. Much remains to be learned about landslide recovery and ecology in the Peace River Region, and the Beatton River landslide could provide an outdoor laboratory for restoration experimentation and trials. Because the landslide is so young and much of the surface is steep and unstable, there are many opportunities to try different slope stabilisation and revegetation measures. The trials could be easily monitored, as access to the landslide is good. Further research should focus on a variety of landslide types, sizes, and ages in the region, and these methods could also be extended to other regions. Similar vegetation and microsite sampling for both the landslide and the surrounding terrain should be employed, preferably with an increased sample size for the relevés and transects and greater utilisation of remotesensed imagery. To better understand the dynamics of colonisation by native plant species, an attempt should be made to locate remote landslides free from the influence of invasive exotic species. Although relatively sparse at the regional scale and generally small in size, landslide ponds may have important implications for landslide stability and ecosystem health. The presence of ponds indicates saturation, therefore pond-bearing landslides near priority management 214 areas should be monitored so that they can be stabilised quickly if they reactivate. These ponds should also be conserved for their ecological value for amphibians and invertebrates, as well as for habitat and a feeding and drinking source for waterfowl and fur-bearing mammals. Further research is needed to investigate the ecological significance of these features. 5.3 Final thoughts The research that formed the basis of this dissertation was initiated to address the gap in knowledge about biophysical diversity and recovery on landslides in northeastern British Columbia’s Peace River Region, a glaciated area highly susceptible to landslide activity. The work is the first of its kind in the region, as there were no previous published diversity studies that collected and analysed detailed field data on landslide vegetation species or ponds, let alone at such a large scale of study. This research provided quantitative confirmation of the increased diversity of some biophysical aspects of landslides compared to undisturbed terrain in northeastern British Columbia and demonstrated that both vegetation and environmental diversity vary with age and size of landslides. The results suggest landslide diversity and recovery evolve over time and may take decades to settle out, and both invasion by exotic vegetation and reactivation of the slide can hinder ecological succession. The findings obtained from the vegetation surveys in this study were used to present recommendations on native plant species to use for restoration of landslides in the Peace River Region and other similar areas. This study also provided valuable baseline information about ponds on landslides, a topic that warrants further investigation given the ecological value of networks of small ponds. Overall, this research has significantly enhanced understanding of landslide diversity and recovery in the 215 Peace River Region and provided timely recommendations for restoration and management of landslides both locally and for similar glaciolacustrine sites around the world. 216 APPENDICES 217 Appendix 1 Material Origin classes The Material Origin variable for the BEC 50m2 plots is a new variable created from components of the soil and site description data collected during BEC sampling. It is intended to describe the primary level of soil development present on each plot site. The coding separates out those plots which have intermediate soil development, as well as those plots which are ponds. Coding is as follows: 1. Mature in situ (benchmark plot in surrounding terrain) Category 1 consists of intact material with a well-developed soil profile, situated in the surrounding terrain. Soils have at least a B horizon, and an A horizon may also be present. In Dystric Brunisols, the A horizon is commonly absent. 2. Mature raft Category 2 consists of mature intact material transported from the surrounding terrain onto the landslide through slope movement processes. The soil is well-developed and has at least a B horizon. An A horizon may also be present. 3. Intermediate development Category 3 consists of partially-developed landslide body soils with an immature B horizon and no A horizon. The B horizon is thin, usually less than 10 cm in thickness. 4. Orthic Regosol Category 4 consists of material on the landslide body which has only a C horizon exposed. The A/B horizons have either been buried, stripped away by movement of material downslope, or eroded away by weathering. 5. Pond Category 5 consists of soils that are inundated under various ponds on the landslide body. Soil development at present is mostly arrested, with soils likely comprised of just a C horizon at the time of flooding. 218 Appendix 2 Vegetation summary tables - Mean species cover (% of total relevé area) Beatton River Landslide Relevés BErl1, BErl2, BErl3 Mean cover SD (%) (+/-) Melilotus officinalis Yellow sweet-clover 28.833 44.312 Equisetum arvense Common horsetail 12.167 19.776 Melilotus alba White sweet-clover 6.000 3.500 Sonchus arvensis Perennial sow-thistle 4.100 2.456 Artemisia sp. 2 Sage sp. 1.767 0.874 Lactuca serriola Prickly lettuce 1.410 1.417 Rubus idaeus Red raspberry 1.087 1.874 Aster ciliolatus Fringed aster 0.700 1.127 Solidago canadensis Canada goldenrod 0.650 0.589 Taraxacum officinale Common dandelion 0.443 0.501 Fragaria virginiana Wild strawberry 0.400 0.529 Cornus stolonifera Red-osier dogwood 0.283 0.407 Unknown forb (white) Forb sp. 0.167 0.289 Brachythecium sp. Ragged moss sp. 0.167 0.289 Populus balsamifera ssp. balsamifera Black cottonwood 0.167 0.208 Chenopodium album Lamb's-quarters 0.133 0.115 Rosa acicularis Prickly rose 0.133 0.153 Achillea millefolium Yarrow 0.123 0.155 Tragopogon dubius Goat's-beard, Yellow salsify 0.103 0.095 Hieracium triste Wooly hawkweed 0.100 0.100 Elymus repens (aka Agropyron repens ) Quackgrass, couch grass 0.100 0.173 Aster sp. Aster sp. 0.083 0.144 Vicia americana American vetch 0.073 0.110 Ranunculus sp. Buttercup sp. 0.067 0.115 Salix sp. 2 Willow sp. 2 0.067 0.115 Salix sp. Willow sp. 0.060 0.053 Shepherdia canadensis Soopolallie 0.043 0.051 Epilobium angustifolium Fireweed 0.037 0.055 Lathyrus ochroleucus Creamy peavine 0.033 0.058 Medicago sativa Alfalfa 0.033 0.058 Elaeagnus commutata Wolf-willow 0.033 0.058 Salix sp. 1 Willow sp. 1 0.033 0.058 Picea glauca White spruce 0.033 0.058 Cirsium arvense Canada thistle 0.013 0.006 Species Common name(s) 219 Beatton River Landslide Relevés BErl1, BErl2, BErl3 -cont'd Species Common name(s) Elymus glaucus Elymus lanceolatus ssp. lanceolatus Lily sp. Trifolium hybridum Bromus inermis ssp. inermis Elymus trachycaulus ssp. trachycaulus Ribes lacustre Blue wildrye Thickspike wheatgrass Lily sp. Alsike clover Smooth brome Slender wheatgrass Black gooseberry 220 Mean cover (%) 0.010 0.010 0.003 0.003 0.003 0.003 0.003 SD (+/-) 0.017 0.017 0.006 0.006 0.006 0.006 0.006 Beatton River Undisturbed Relevés BEru1, BEru2d, BEru3 Species Common name(s) Amelanchier alnifolia Betula papyrifera Aralia nudicaulis Linnaea borealis Carex sp. 1 Viburnum edule Rosa acicularis Picea glauca Carex sp. 3 Aster conspicuus Alnus viridis ssp. sinuata Cornus stolonifera Leymus innovatus Calamagrostis canadensis Shepherdia canadensis Symphoricarpos albus Apocynum androsaemifolium Salix sp. 2 Artemisia sp. 1 Populus tremuloides Rubus pubescens Elaeagnus commutata Tragopogon dubius Hesperostipa curtiseta Pyrola asarifolia Prunus virginiana Lonicera dioica Populus balsamifera ssp. balsamifera Spiraea betulifolia Lathyrus ochroleucus Cornus canadensis Platydictya jungermannioides Vicia americana Saskatoon Paper birch Wild sarsaparilla Twinflower Sedge sp. 1 Highbush-cranberry Prickly rose White spruce Sedge sp. 3 Showy aster Sitka alder Red-osier dogwood Fuzzy-spiked wildrye Bluejoint Soopolallie Common snowberry Spreading dogbane Willow sp. 2 Sage sp. 1 Trembling aspen Trailing raspberry Wolf-willow Goat's-beard, yellow salsify Needle-and-thread grass Pink wintergreen Choke cherry Red honeysuckle Black cottonwood Birch-leaved spirea Creamy peavine Bunchberry False willow moss, Spruce's leskea American vetch 221 Mean cover SD (%) (+/-) 10.846 16.598 10.667 18.475 7.824 2.299 6.832 6.135 5.667 9.815 5.550 4.877 4.925 0.393 4.727 6.724 4.000 6.928 3.852 3.372 3.500 6.062 2.683 2.675 2.625 2.770 2.614 3.105 2.407 3.049 2.003 3.038 1.833 3.175 1.510 2.615 1.417 2.454 1.297 1.101 1.268 1.874 1.083 1.876 1.000 1.732 0.667 1.155 0.500 0.621 0.422 0.298 0.396 0.487 0.396 0.351 0.333 0.577 0.329 0.333 0.277 0.392 0.233 0.400 0.233 0.354 Beatton River Undisturbed Relevés BEru1, BEru2d, BEru3 - cont'd Mean cover SD (%) (+/-) Galium boreale Northern bedstraw 0.213 0.191 Mitella nuda Common mitrewort 0.169 0.291 Maianthemum canadense Wild lily-of-the-valley 0.168 0.148 "Silver grass" Grass sp. 0.167 0.289 Fragaria vesca Wood strawberry 0.165 0.188 Salix sp. 3 Willow sp. 3 0.162 0.193 Disporum trachycarpum Rough-fruited fairybells 0.149 0.157 Ribes lacustre Black gooseberry 0.148 0.129 Mertensia paniculata Tall bluebell 0.143 0.128 Equisetum arvense Common horsetail 0.127 0.217 Ribes triste Red swamp currant 0.125 0.110 Actaea rubra Baneberry 0.100 0.163 Orthilia secunda One-sided wintergreen 0.095 0.095 Fragaria virginiana Wild strawberry 0.094 0.126 Epilobium angustifolium Fireweed 0.093 0.089 Comandra umbellata Bastard toadflax 0.093 0.162 Aster ciliolatus Fringed aster 0.083 0.018 Achnatherum nelsonii Columbia needlegrass 0.083 0.144 Hesperostipa spartea ( aka Stipa spartea) Porcupinegrass 0.083 0.144 Peltigera sp. Pelt lichen sp. 0.080 0.122 Rubus idaeus Red raspberry 0.079 0.097 Lonicera involucrata Black twinberry 0.073 0.082 Osmorhiza chilensis Mountain sweet-cicely 0.068 0.117 Hylocomium splendens Step moss 0.066 0.095 Plagiomnium cuspidatum Baby tooth moss, woodsy thyme-moss 0.056 0.048 Maianthemum canadense Violet sp. 0.055 0.048 Ribes oxyacanthoides Northern gooseberry 0.048 0.048 Eurynchiastrum pulchellum Elegant feather-moss 0.040 0.037 Galium triflorum Sweet-scented bedstraw 0.040 0.038 Achillea millefolium Yarrow 0.039 0.053 Geocaulon lividum False toadflax, northern comandra 0.033 0.058 Sonchus sp. Sow-thistle sp. 0.033 0.058 Androsace septentrionalis Pygmyflower rockjasmine 0.033 0.058 Species Common name(s) 222 Beatton River Undisturbed Relevés BEru1, BEru2d, BEru3 - cont'd Species Common name(s) Ribes sp. Smilacina stellata Taraxacum officinale Erigeron glabellus Koelaria macrantha Pleurozium schreberi Pohlia nutans Pyrola sp. Erigeron acris Agropyron sp. Syntrichia ruralis Solidago canadensis Salix sp. 1 Trifolium pratense Petasites frigidus Dicranum polysetum Gymnocarpium dryopteris Delphinium sp. Dryas drummondii Maianthemum stellatum Rhinanthus minor Glyceria sp. Platanthera dilitata Cladonia sp. Juniperus communis Gooseberry/currant sp. Star-flowered false Solomon's seal Common dandelion Smooth fleabane Junegrass Red-stemmed feathermoss Nodding thread-moss Wintergreen sp. Bitter fleabane Slender wheatgrass Twisted moss Canada goldenrod Willow sp. 1 Clover sp. Sweet coltsfoot Broom moss, wavyleaf moss Oak fern Larkspur sp. Yellow mountain avens Starry false lily of the valley Yellow rattle Manna grass sp. White northern bog-orchid Pixie cup sp. Common juniper 223 Mean cover SD (%) (+/-) 0.033 0.058 0.023 0.030 0.018 0.018 0.017 0.029 0.017 0.029 0.014 0.017 0.013 0.022 0.012 0.014 0.010 0.017 0.010 0.017 0.008 0.013 0.008 0.014 0.008 0.013 0.007 0.012 0.004 0.007 0.003 0.004 0.003 0.004 0.003 0.006 0.003 0.006 0.003 0.006 0.003 0.006 0.003 0.006 0.002 0.003 0.002 0.003 0.002 0.003 Cecil Lake Landslide Relevés CErl1, CErl2, CErl3 Species Common name(s) Equisetum arvense Common horsetail Alnus viridis ssp. sinuata Sitka alder Petasites frigidus Sweet coltsfoot Salix sp. 1 Willow sp. 1 Populus balsamifera ssp. balsamifera Black cottonwood Shepherdia canadensis Soopolallie Picea glauca White spruce Populus tremuloides Trembling aspen Rosa acicularis Prickly rose Salix sp. 2 Willow sp. 2 Trifolium hybridum Alsike clover Salix sp. 3 Willow sp. 3 Viburnum edule Highbush-cranberry Cornus stolonifera Red-osier dogwood Calamagrostis canadensis Bluejoint Taraxacum officinale Common dandelion Typha sp. Cattail sp. Salix sp. 6 Willow sp. 6 Eurynchiastrum pulchellum Elegant feather-moss Rubus idaeus Red raspberry Aster ciliolatus Fringed aster Epilobium angustifolium Fireweed Betula papyrifera Paper birch Cirsium arvense Canada thistle Linnaea borealis Twinflower Fragaria virginiana Wild strawberry Sonchus arvensis Perennial sow-thistle Rubus pubescens Trailing raspberry Vicia americana American vetch Salix sp. 4 Willow sp. 4 Lathyrus ochroleucus Creamy peavine Aralia nudicaulis Wild sarsaparilla Pyrola asarifolia Pink wintergreen 224 Mean cover SD (%) (+/-) 33.500 12.173 24.092 20.687 6.733 11.490 5.630 4.352 4.767 6.280 3.128 0.327 2.679 2.065 2.642 2.455 2.412 0.390 1.958 2.895 1.664 2.516 1.518 1.885 1.492 1.784 1.325 0.282 1.258 1.906 0.833 0.441 0.675 0.622 0.667 1.155 0.567 0.787 0.523 0.285 0.518 0.729 0.415 0.388 0.350 0.563 0.348 0.410 0.332 0.282 0.329 0.170 0.298 0.495 0.265 0.190 0.253 0.137 0.225 0.390 0.203 0.086 0.197 0.341 0.194 0.319 Cecil Lake Landslide Relevés CErl1, CErl2, CErl3 - cont'd Species Aster conspicuus Brachythecium sp. 2 Salix sp. 5 Solidago canadensis Amelanchier alnifolia Mertensia paniculata Ribes oxyacanthoides Cornus canadensis Phleum pratense Elymus trachycaulus Brachythecium sp. 1 Salix exigua Hordeum jubatum Symphoricarpos albus Delphinium glaucum Lonicera involucrata Goodyera repens "Hairy grass" Brachythecium sp. 3 Pohlia nutans Cinna latifolia Galium triflorum Fragaria vesca Glyceria sp. Achillea sibirica Melilotus officinalis Bromus inermis ssp. inermis "Tallwheat grass" sp. Achillea millefolium Ribes sp. Geum macrophyllum Galium boreale Matricaria perforata Mean cover (%) Showy aster 0.170 Ragged moss sp. 0.167 Willow sp. 5 0.167 Canada goldenrod 0.161 Saskatoon 0.117 Tall bluebell 0.108 Northern gooseberry 0.092 Bunchberry 0.091 Timothy 0.090 Slender wheat grass 0.084 Moss sp. 0.067 Sandbar willow 0.067 Foxtail barley 0.064 Common snowberry 0.060 Tall larkspur 0.057 Black twinberry 0.057 Dwarf rattlesnake-plantain 0.050 "Hairy" grass sp. 0.049 Moss sp. 0.039 Nodding thread-moss 0.037 Nodding wood-reed 0.033 Sweet-scented bedstraw 0.026 Wood strawberry 0.024 Manna grass sp. 1 0.021 Siberian yarrow 0.020 Yellow sweet-clover 0.019 Smooth brome 0.019 "Tallwheat" grass sp. 0.018 Common yarrow 0.016 Gooseberry/currant sp. 0.016 Large-leaved avens 0.015 Northern bedstraw 0.008 Scentless chamomile 0.008 Common name(s) 225 SD (+/-) 0.255 0.289 0.289 0.177 0.126 0.091 0.103 0.077 0.087 0.146 0.115 0.115 0.051 0.100 0.098 0.068 0.087 0.085 0.068 0.064 0.058 0.017 0.042 0.034 0.026 0.033 0.033 0.032 0.018 0.027 0.022 0.010 0.014 Cecil Lake Landslide Relevés CErl1, CErl2, CErl3 - cont'd Species Ribes triste Dicranum sp. Funaria sp. Epilobium ciliatum Mitella nuda Spiraea sp. Agrostis stolonifera Cladonia sp. Nephroma sp. Lonicera dioica Bromus ciliatus Poa nemoralis ssp. interior Elymus lanceolatus ssp. lanceolatus Peltigera sp. Actaea rubra Hylocomium splendens Achillea sp. Chenopodium sp. Crepis tectorum Leucantheum sp. Sanicula marilandica Koelaria macrantha Rush sp. Ribes sp. 2 Ribes lacustre Rumex crispus Platanthera dilitata Viola sp. Poa pratensis Ribes laxiflorum Marchantia sp. "Bottlebrush moss" sp. Pleurozium schreberi Mean cover (%) Red swamp currant 0.008 Broom moss sp. 0.007 Rope moss sp. 0.007 Purple-leaved willowherb 0.007 Common mitrewort 0.007 Hardhack sp. 0.007 Redtop 0.007 Pixie cup sp. 0.007 Kidney lichen sp. 0.007 Red honeysuckle 0.006 Fringed brome 0.005 Inland blue grass 0.005 Thickspike wheatgrass 0.005 Pelt lichen sp. 0.005 Baneberry 0.004 Step moss 0.003 Yarrow sp. 0.003 Goosefoot sp. 0.003 Annual hawksbeard 0.003 Daisy sp. 0.003 Maryland black snakeroot 0.003 Junegrass 0.003 Rush sp. 0.003 Gooseberry/currant sp. 2 0.003 Black gooseberry 0.003 Curly dock 0.002 White northern bog-orchid 0.002 Violet sp. 0.002 Kentucky bluegrass 0.002 Trailing black currant 0.002 Liverwort sp. 0.001 "Bottlebrush" moss sp. 0.001 Red-stemmed feathermoss 0.001 Common name(s) 226 SD (+/-) 0.014 0.012 0.012 0.012 0.012 0.012 0.012 0.012 0.012 0.010 0.009 0.009 0.009 0.009 0.007 0.006 0.006 0.006 0.006 0.006 0.004 0.006 0.004 0.006 0.004 0.003 0.003 0.001 0.003 0.003 0.001 0.001 0.001 Cecil Lake Landslide Relevés CErl1, CErl2, CErl3 - cont'd Species Common name(s) Hieracium triste Senecio eremophilus Sonchus arvensis Gentianella amarella "Bigwheat grass" sp. Glyceria sp. 2 Beckmannia syzigachne Wooly hawkweed Cutleaf groundsel Perennial sow thistle Autumn gentian "Bigwheat" grass sp. Manna grass sp. 2 American sloughgrass 227 Mean cover (%) 0.001 0.001 0.001 0.001 0.001 0.001 0.001 SD (+/-) 0.001 0.001 0.001 0.001 0.001 0.001 0.001 Cecil Lake Undisturbed Relevés CEru1, CEru2a, CEru3 Species Common name(s) Picea glauca Populus tremuloides Rosa acicularis Viburnum edule Aralia nudicaulis Hylocomium splendens Shepherdia canadensis Alnus viridis ssp. sinuata Betula papyrifera Linnaea borealis "Fuzzy grass" sp. Cornus stolonifera Pleurozium schreberi Aster conspicuus Cornus canadensis Amelanchier alnifolia Rubus pubescens Ribes triste Symphoricarpos albus Mertensia paniculata Ptilium crista-castrensis Ribes lacustre Lonicera dioica "Thin grass" sp. Calamagrostis canadensis Salix sp. Lathyrus ochroleucus Clematis sp. Lonicera involucrata Rhytidiadelphus triquetrus Mitella nuda Eurynchiastrum pulcellum Disporum trachycarpum White spruce Trembling aspen Prickly rose Highbush-cranberry Wild sarsaparilla Step moss Soopolallie Sitka alder Paper birch Twinflower "Fuzzy" grass sp. Red-osier dogwood Red-stemmed feathermoss Showy aster Bunchberry Saskatoon Trailing raspberry Red swamp currant Common snowberry Tall bluebell Knight's plume Black gooseberry Red honeysuckle "Thin" grass sp. Bluejoint Willow sp. Creamy peavine Clematis sp. Black twinberry Electrified cat's-tail moss Common mitrewort Elegant beaked moss Rough-fruited fairybells 228 Mean cover SD (%) (+/-) 22.893 14.687 9.107 12.483 6.908 3.425 6.762 3.772 6.460 1.935 6.210 5.743 4.388 4.020 4.015 2.953 3.624 5.017 3.137 3.102 2.253 3.894 1.922 1.738 1.118 1.779 1.118 0.986 1.058 0.820 0.971 0.806 0.782 0.500 0.445 0.404 0.426 0.224 0.349 0.327 0.323 0.543 0.297 0.285 0.279 0.106 0.253 0.439 0.240 0.235 0.190 0.329 0.146 0.156 0.133 0.231 0.115 0.133 0.110 0.191 0.108 0.166 0.087 0.150 0.086 0.012 Cecil Lake Undisturbed Relevés CEru1, CEru2a, CEru3 - cont'd Mean cover (%) Orthilia secunda One-sided wintergreen 0.081 Geocaulon lividum Bastard toad-flax 0.060 Aster ciliolatus Fringed aster 0.053 Galium boreale Northern bedstraw 0.053 Pyrola chlorantha Green wintergreen 0.047 Peltigera polydactylon Pioneer pelt 0.047 Pyrola asarifolia Pink wintergreen 0.038 Ribes oxyacanthoides Northern gooseberry 0.035 Fragaria virginiana Wild strawberry 0.033 Rubus idaeus Red raspberry 0.029 Brachythecium sp. Ragged moss sp. 0.027 Fragaria vesca Wood strawberry 0.027 Moss sp. Moss sp. 0.020 Fern sp. Fern sp. 0.020 Epilobium angustifolium Fireweed 0.020 Actaea rubra Baneberry 0.018 Osmorhiza chilensis Mountain sweet-cicely 0.018 Dicranum polysetum Wavy leaf moss, broom moss 0.014 Galium triflorum Sweet-scented bedstraw 0.012 Dicranum scoparium Broom forkmoss 0.010 Plagiomnium cuspidatum Baby tooth moss, woodsy thyme moss 0.007 Viola sp. Violet sp. 0.007 Vicia americana American vetch 0.005 Delphinium glaucum Tall larkspur 0.004 Equisetum arvense Common horsetail 0.003 Goodyera oblongifolia Rattlesnake-plantain 0.003 Listera sp. (aka Neottia sp.) Twayblade sp. 0.003 Maianthemum canadense Wild lily-of-the-valley 0.003 Taraxacum officinale Common dandelion 0.003 Cladonia sp. Pixie cup sp. 0.003 Prunus virginiana Choke cherry 0.003 Gymnocarpium dryopteris Oak fern 0.002 Species Common name(s) 229 SD (+/-) 0.059 0.060 0.084 0.051 0.072 0.045 0.013 0.061 0.058 0.026 0.046 0.025 0.035 0.035 0.035 0.011 0.028 0.025 0.013 0.017 0.012 0.012 0.005 0.007 0.006 0.006 0.006 0.006 0.006 0.006 0.006 0.003 Hasler Flats Landslide Relevés HArl1b, HArl2b, HArl3a Species Common name(s) Equisetum arvense Populus tremuloides Rosa acicularis Alnus viridis ssp. sinuata Rubus idaeus Cornus stolonifera Symphoricarpos albus Pohlia nutans Viburnum edule Lonicera dioica Smilacina racemosa Amelanchier alnifolia Aralia nudicaulis Prunus virginiana Calamagrostis canadensis Bromus inermis Aster conspicuus Lonicera involucrata Elymus glaucus ssp. glaucus Ribes oxyacanthoides Salix sp. 2 Rubus pubescens Ribes triste Populus balsamifera ssp. balsamifera Mertensia paniculata Salix sp. 1 Petasites frigidus Vicia americana Streptopus amplexifolius Viola sp. Marchantia polymorpha Aster ciliolatus Lathyrus ochroleucus Common horsetail Trembling aspen Prickly rose Sitka alder Red raspberry Red-osier dogwood Common snowberry Nodding thread-moss Highbush-cranberry Red honeysuckle False Solomon's-seal Saskatoon Wild sarsaparilla Choke cherry Bluejoint Smooth brome Showy aster Black twinberry Blue wildrye Northern gooseberry Willow sp. 2 Trailing raspberry Red swamp currant Black cottonwood Tall bluebell Willow sp. 1 Sweet coltsfoot American vetch Clasping twistedstalk Violet sp. Common liverwort Fringed aster Creamy peavine 230 Mean cover (%) 16.417 10.758 10.375 6.308 5.595 4.900 4.145 2.423 2.423 2.348 1.623 1.528 1.403 1.320 1.088 1.051 0.925 0.716 0.696 0.461 0.434 0.348 0.335 0.300 0.281 0.266 0.248 0.247 0.232 0.199 0.198 0.176 0.176 SD (+/-) 1.041 5.329 2.211 4.242 3.055 3.776 2.427 0.505 1.273 2.706 2.537 1.671 2.147 0.437 0.934 1.818 0.839 0.584 0.654 0.491 0.475 0.378 0.273 0.173 0.110 0.142 0.161 0.178 0.401 0.313 0.208 0.172 0.204 Hasler Flats Landslide Relevés HArl1b, HArl2b, HArl3a - cont'd Species Common name(s) Galium boreale Northern bedstraw Betula papyrifera Paper birch Heracleum lanatum Cow-parsnip Actaea rubra Baneberry Smilacina stellata Star-flowered false Solomon's-seal Taraxacum officinale Common dandelion Fragaria vesca Wood strawberry Epilobium angustifolium Fireweed Cornus canadensis Bunchberry Mitella nuda Common mitrewort Galium triflorum Sweet-scented bedstraw Picea glauca White spruce Dicranum polysetum Wavy leaf moss, broom moss Pyrola asarifolia Pink wintergreen Thalictrum occidentale Western meadowrue Fragaria virginiana Wild strawberry Cirsium arvense Canada thistle Spiraea betulifolia Birch-leaved spirea Maianthemum canadense Wild lily-of-the-valley Poa palustris Fowl blue grass Sonchus arvensis Perennial sow-thistle Epilobium ciliatum Purple-leaved willowherb Linnaea borealis Twinflower Veratrum viride Indian hellebore "Oat grass" sp. "Oat grass" grass sp. Rubus parviflorus Thimbleberry Bromus ciliatus Fringed brome Typha sp. Cattail sp. Leymus innovatus Fuzzy-spiked wildrye Veronica beccabunga European speedwell, brooklime Elymus trachycaulus ssp. trachycaulus Slender wheatgrass Carex sp. Sedge sp. Carex rostrata Beaked sedge 231 Mean cover (%) 0.163 0.103 0.098 0.090 0.088 0.088 0.084 0.082 0.074 0.068 0.065 0.063 0.062 0.061 0.054 0.047 0.046 0.040 0.039 0.038 0.030 0.029 0.027 0.023 0.019 0.019 0.016 0.015 0.012 0.010 0.010 0.010 0.008 SD (+/-) 0.142 0.003 0.100 0.089 0.153 0.052 0.135 0.085 0.004 0.089 0.061 0.097 0.107 0.093 0.019 0.081 0.058 0.034 0.046 0.026 0.014 0.020 0.031 0.040 0.033 0.019 0.027 0.020 0.020 0.017 0.011 0.017 0.009 Hasler Flats Landslide Relevés HArl1b, HArl2b, HArl3a - cont'd Species Common name(s) Cirsium vulgare Festuca sp. Achillea millefolium Trisetum cernuum "Limp grass" sp. Syntrichia ruralis Plagiomnium cuspidatum Cyclamen hederifolium Hieracium sabaudum Orthilia secunda "Frothy grass" sp. Shepherdia canadensis Leptobryum pyriforme Fern sp. Equisetum variegatum Geum macrophyllum Chenopodium capitatum Boechera stricta Plantago major "Pondweed" sp. Trifolium hybridum Phleum pratense Ribes hudsonianum Ribes laxiflorum Bull thistle Fescue sp. Yarrow Nodding trisetum "Limp grass" grass sp. Twisted moss Baby tooth moss, woodsy thyme-moss Hardy cyclamen, ivy leaved cyclamen European hawkweed, Savoy hawkweed One-sided wintergreen "Frothy grass" grass sp. Soopolallie Golden thread-moss Fern sp. Variegated scouring-rush Large-leaved avens Strawberry-blite Canada rockcress Broadleaf plantain Pondweed sp. Alsike clover Timothy Northern black currant Trailing black currant 232 Mean cover (%) 0.007 0.007 0.005 0.005 0.005 0.004 0.004 0.003 0.003 0.003 0.003 0.003 0.002 0.002 0.002 0.002 0.001 0.001 0.001 0.001 0.001 0.001 0.001 0.001 SD (+/-) 0.012 0.012 0.005 0.009 0.009 0.007 0.007 0.006 0.006 0.004 0.004 0.004 0.003 0.003 0.003 0.003 0.001 0.001 0.001 0.001 0.001 0.001 0.001 0.001 Hasler Flats Undisturbed Relevés HAru1, HAru2a, HAru3a Species Common name(s) Equisetum arvense Populus tremuloides Alnus viridis ssp. sinuata Cornus stolonifera Viburnum edule Aralia nudicaulis Symphoricarpos albus Rosa acicularis Populus balsamifera ssp. balsamifera Heracleum lanatum Aster conspicuus Amelanchier alnifolia Lonicera dioica Prunus virginiana Mitella nuda Rubus idaeus Salix sp. 2 Ribes oxyacanthoides Smilacina racemosa Actaea rubra Rubus pubescens Thalictrum occidentale Picea glauca Smilacina stellata Ribes triste Lonicera involucrata Viola sp. Lathyrus ochroleucus Maianthemum canadense Streptopus amplexifolius Vicia americana Petasites frigidus Angelica lucida Common horsetail Trembling aspen Sitka alder Red-osier dogwood Highbush-cranberry Wild sarsaparilla Common snowberry Prickly rose Black cottonwood Cow-parsnip Showy aster Saskatoon Red honeysuckle Choke cherry Common mitrewort Red raspberry Willow sp. 2 Northern gooseberry False Solomon's-seal Baneberry Trailing raspberry Western meadowrue White spruce Star-flowered false Solomon's-seal Red swamp currant Black twinberry Violet sp. Creamy peavine Wild lily-of-the-valley Clasping twistedstalk American vetch Sweet coltsfoot Seacoast angelica, sea-watch 233 Mean cover SD (%) (+/-) 15.625 6.945 13.580 5.074 9.863 17.084 7.358 4.376 6.983 5.925 6.856 4.865 5.378 5.090 5.283 1.617 4.335 6.321 3.844 5.059 2.556 2.311 2.503 1.616 1.976 2.900 1.609 1.262 1.397 1.984 1.236 1.720 1.026 0.876 0.904 1.521 0.653 0.481 0.580 0.544 0.551 0.240 0.516 0.532 0.510 0.515 0.488 0.420 0.487 0.318 0.438 0.193 0.415 0.595 0.328 0.285 0.327 0.375 0.276 0.386 0.235 0.147 0.231 0.138 0.199 0.179 Hasler Flats Undisturbed Relevés HAru1, HAru2a, HAru3a - cont'd Species Salix sp. 1 Calamagrostis canadensis Pyrola asarifolia Aster foliaceus Ribes hudsonianum Mertensia paniculata Cirsium arvense Aster ciliolatus Dicranum polysetum Galium boreale Shepherdia canadensis Galium triflorum "Mystery grass 2" sp. Disporum trachycarpum Plagiomnium cuspidatum Fragaria vesca Plagiomnium ellipticum Osmorhiza chilensis Cornus canadensis Salix sp. 3 Acer glabrum Orthilia secunda Bromus inermis ssp. inermis Fragaria virginiana Taraxacum officinale "Hairy grass" Epilobium ciliatum Veronica beccabunga Carex disperma "Mystery grass" sp. 1 Veratrum viride Linnaea borealis Bromus ciliatus Mean cover (%) Willow sp. 1 0.195 Bluejoint 0.185 Pink wintergreen 0.163 Leafy-bracted aster 0.131 Northern black currant 0.130 Tall bluebell 0.124 Creeping thistle 0.123 Fringed aster 0.110 Wavy leaf moss, broom moss 0.105 Northern bedstraw 0.085 Soopolallie 0.083 Sweet-scented bedstraw 0.077 Mystery grass sp. 2 0.077 Rough-fruited fairybells 0.074 Baby tooth moss, woodsy thyme-moss 0.068 Wood strawberry 0.056 Marsh thyme-moss 0.041 Mountain sweet-cicely 0.035 Bunchberry 0.034 Willow sp. 3 0.033 Douglas maple 0.033 One-sided wintergreen 0.032 Smooth brome 0.028 Wild strawberry 0.027 Common dandelion 0.026 "Hairy grass" grass sp. 0.024 Purple-leaved willowherb 0.019 European speedwell, brooklime 0.018 Soft-leaf sedge, two-seed sedge 0.018 "Mystery grass" grass sp. 1 0.017 Indian hellebore 0.014 Twinflower 0.011 Fringed brome 0.008 Common name(s) 234 SD (+/-) 0.287 0.215 0.249 0.218 0.225 0.027 0.214 0.132 0.173 0.071 0.144 0.028 0.133 0.128 0.118 0.097 0.056 0.044 0.049 0.058 0.058 0.051 0.018 0.031 0.030 0.042 0.033 0.030 0.016 0.027 0.025 0.019 0.014 Hasler Flats Undisturbed Relevés HAru1, HAru2a, HAru3a - cont'd Species Common name(s) Poa pratensis Equisetum pratense Spiraea betulifolia Leymus innovatus Elymus lanceolatus ssp. lanceolatus Epilobium angustifolium Geum rivale Phleum pratense "Soft grass" sp Grass sp. Carex sp. Rubus parviflorus Botrychium virginianum Cyclamen hederifolium Glyceria sp. Achillea millefolium Monotropa uniflora Ribes lacustre Kentucky bluegrass Meadow horsetail, shade horsetail Birch-leaved spirea Fuzzy-spiked wildrye Thickspike wheatgrass Fireweed Water avens, purple avens Timothy "Soft grass" grass sp. Unknown grass sp. Sedge sp. Thimbleberry Rattlesnake fern Hardy cyclamen, ivy leaved cyclamen Manna grass sp. Yarrow Ghost pipe, Indian-pipe Black gooseberry 235 Mean cover (%) 0.008 0.007 0.005 0.005 0.005 0.003 0.003 0.003 0.003 0.003 0.003 0.003 0.002 0.002 0.002 0.001 0.001 0.001 SD (+/-) 0.014 0.006 0.009 0.009 0.004 0.003 0.006 0.004 0.004 0.006 0.006 0.004 0.003 0.003 0.003 0.001 0.001 0.001 Appendix 3 Vegetation cover by growth form Beatton River Landslide Relevés - Total mean vegetation cover = 59.68% Species Trees Populus balsamifera ssp. balsamifera Picea glauca Mean cover (%) Total Proportion of all cover 0.167 0.033 0.200 0.335 Total Proportion of all cover 1.087 0.283 0.133 0.067 0.060 0.043 0.033 0.033 0.003 1.742 2.919 Shrubs Rubus idaeus Cornus stolonifera Rosa acicularis Salix sp. 2 Salix sp. Shepherdia canadensis Elaeagnus commutata Salix sp. 1 Ribes lacustre Forbs Melilotus officinalis Melilotus alba Sonchus arvensis Artemisia sp. 2 Lactuca serriola Aster ciliolatus Solidago canadensis Taraxacum officinale Fragaria virginiana Unknown forb (white) Chenopodium album Achillea millefolium Tragopogon dubius Hieracium triste Aster sp. Vicia americana Ranunculus sp. 28.833 6.000 4.100 1.767 1.410 0.700 0.650 0.443 0.400 0.167 0.133 0.123 0.103 0.100 0.083 0.073 0.067 236 Beatton River Landslide Relevés - cont'd Forbs -cont'd Epilobium angustifolium Lathyrus ochroleucus Medicago sativa Cirsium arvense Lily sp. Trifolium hybridum Total Proportion of all cover Graminoids Elymus repens (aka Agropyron repens ) Elymus glaucus Elymus lanceolatus ssp. lanceolatus Bromus inermis ssp. inermis Elymus trachycaulus ssp. trachycaulus 0.037 0.033 0.033 0.013 0.003 0.003 45.274 75.861 Total Proportion of all cover 0.100 0.010 0.010 0.003 0.003 0.126 0.211 Total Proportion of all cover 12.167 12.167 20.387 Total Proportion of all cover 0.167 0.167 0.280 Ferns & Fern allies Equisetum arvense Bryophytes Brachythecium sp. 237 Beatton River Undisturbed Relevés - Total mean vegetation cover = 97.348% Species Trees Betula papyrifera Picea glauca Populus tremuloides Populus balsamifera ssp. balsamifera Mean cover (%) Total Proportion of all cover 10.667 4.727 1.297 0.396 17.087 17.552 Total Proportion of all cover 10.846 5.550 4.925 3.500 2.683 2.407 2.003 1.510 1.268 1.083 0.422 0.396 0.162 0.148 0.125 0.079 0.073 0.048 0.033 0.008 0.002 37.271 38.286 Shrubs Amelanchier alnifolia Viburnum edule Rosa acicularis Alnus viridis ssp. sinuata Cornus stolonifera Shepherdia canadensis Symphoricarpos albus Salix sp. 2 Rubus pubescens Elaeagnus commutata Prunus virginiana Lonicera dioica Salix sp. Ribes lacustre Ribes triste Rubus idaeus Lonicera involucrata Ribes oxyacanthoides Ribes sp. Salix sp. 1 Juniperus communis Forbs Aralia nudicaulis Linnaea borealis Aster conspicuus 7.824 6.832 3.852 238 Beatton River Undisturbed Relevés - cont'd Species Forbs - cont'd Apocynum androsaemifolium Artemisia sp. 1 Tragopogon dubius Pyrola asarifolia Spiraea betulifolia Lathyrus ochroleucus Cornus canadensis Vicia americana Galium boreale Mitella nuda Maianthemum canadense Fragaria vesca Disporum trachycarpum (aka Prosartes trachycarpa ) Mertensia paniculata Actaea rubra Orthilia secunda Fragaria virginiana Epilobium angustifolium Comandra umbellata Aster ciliolatus Osmorhiza chilensis Maianthemum canadense Galium triflorum Achillea millefolium Geocaulon lividum Sonchus sp. Smilacina stellata Taraxacum officinale Erigeron glabellus Pyrola sp. Erigeron acris Solidago canadensis Trifolium pratense Petasites frigidus Delphinium sp. 239 Mean cover (%) 1.833 1.417 1.000 0.500 0.333 0.329 0.277 0.233 0.213 0.169 0.168 0.165 0.149 0.143 0.100 0.095 0.094 0.093 0.093 0.083 0.068 0.055 0.040 0.039 0.033 0.033 0.023 0.018 0.017 0.012 0.010 0.008 0.007 0.004 0.003 Beatton River Undisturbed Relevés - cont'd Species Forbs - cont'd Dryas drummondii Maianthemum stellatum Rhinanthus minor Platanthera dilitata Mean cover (%) Total Proportion of all cover Graminoids Carex sp. 1 Carex sp. 3 Leymus innovatus Calamagrostis canadensis Hesperostipa curtiseta "Silver grass" Achnatherum nelsonii Hesperostipa spartea (aka Stipa spartea ) Androsace septentrionalis Koelaria macrantha Elymus trachycaulus ssp. trachycaulus Glyceria sp. 0.003 0.003 0.003 0.002 26.376 27.095 Total Proportion of all cover 5.667 4.000 2.625 2.614 0.667 0.167 0.083 0.083 0.033 0.017 0.010 0.003 15.969 16.404 Total Proportion of all cover 0.127 0.003 0.130 0.134 Ferns & Fern allies Equisetum arvense Gymnocarpium dryopteris Bryophytes Platydictya jungermannioides Hylocomium splendens Plagiomnium cuspidatum Eurynchiastrum pulchellum Pleurozium schreberi 0.233 0.066 0.056 0.040 0.014 240 Beatton River Undisturbed Relevés - cont'd Species Bryophytes - cont'd Pohlia nutans Syntrichia ruralis Dicranum polysetum Mean cover (%) Total Proportion of all cover 0.013 0.008 0.003 0.433 0.445 Total Proportion of all cover 0.080 0.002 0.082 0.084 Lichens Peltigera sp. Cladonia sp. 241 Cecil Lake Landslide Relevés - Total mean vegetation cover = 104.160% Species Trees Populus balsamifera ssp. balsamifera Picea glauca Populus tremuloides Betula papyrifera Mean cover (%) Total Proportion of all cover 4.767 2.679 2.642 0.350 10.438 10.021 Total Proportion of all cover 24.092 5.630 3.128 2.412 1.958 1.518 1.492 1.325 0.667 0.523 0.265 0.225 0.167 0.117 0.092 0.067 0.060 0.057 0.016 0.008 0.006 0.003 0.003 0.002 43.833 42.082 Shrubs Alnus viridis ssp. sinuata Salix sp. 1 Shepherdia canadensis Rosa acicularis Salix sp. 2 Salix sp. 3 Viburnum edule Cornus stolonifera Salix sp. 6 Rubus idaeus Rubus pubescens Salix sp. 4 Salix sp. 5 Amelanchier alnifolia Ribes oxyacanthoides Salix exigua Symphoricarpos albus Lonicera involucrata Ribes sp. Ribes triste Lonicera dioica Ribes sp. 2 Ribes lacustre Ribes laxiflorum 242 Cecil Lake Landslide Relevés - cont'd Species Forbs Petasites frigidus Trifolium hybridum Taraxacum officinale Typha sp. Aster ciliolatus Epilobium angustifolium Cirsium arvense Linnaea borealis Fragaria virginiana Sonchus arvensis Vicia americana Lathyrus ochroleucus Aralia nudicaulis Pyrola asarifolia Aster conspicuus Solidago canadensis Mertensia paniculata Cornus canadensis Delphinium glaucum Goodyera repens Galium triflorum Fragaria vesca Achillea sibirica Melilotus officinalis Achillea millefolium Geum macrophyllum Galium boreale Matricaria perforata (aka Tripleurospermum inodorum ) Epilobium ciliatum Mitella nuda Spiraea sp. Actaea rubra Achillea sp. Chenopodium sp. 243 Mean cover (%) 6.733 1.664 0.833 0.675 0.518 0.415 0.348 0.332 0.329 0.298 0.253 0.203 0.197 0.194 0.170 0.161 0.108 0.091 0.057 0.050 0.026 0.024 0.020 0.019 0.016 0.015 0.008 0.008 0.007 0.007 0.007 0.004 0.003 0.003 Cecil Lake Landslide Relevés - cont'd Species Forbs - cont'd Crepis tectorum Leucantheum sp. Sanicula marilandica Rumex crispus Platanthera dilitata Viola sp. Hieracium triste Senecio eremophilus Sonchus arvensis Gentianella amarella Mean cover (%) Total Proportion of all cover Graminoids Calamagrostis canadensis Phleum pratense Elymus trachycaulus Hordeum jubatum "Hairy grass" Cinna latifolia Glyceria sp. Bromus inermis ssp. inermis "Tallwheat grass" Agrostis stolonifera Bromus ciliatus Poa nemoralis ssp. interior Elymus lanceolatus ssp. lanceolatus Koelaria macrantha Rush sp. Poa pratensis "Bigwheat grass" Glyceria sp. 2 Beckmannia syzigachne Total Proportion of all cover 244 0.003 0.003 0.003 0.002 0.002 0.002 0.001 0.001 0.001 0.001 13.815 13.263 1.258 0.090 0.084 0.064 0.049 0.033 0.021 0.019 0.018 0.007 0.005 0.005 0.005 0.003 0.003 0.002 0.001 0.001 0.001 1.669 1.602 Cecil Lake Landslide Relevés - cont'd Species Ferns & Fern allies Equisetum arvense Mean cover (%) Total Proportion of all cover 33.500 33.500 32.162 Total Proportion of all cover 0.567 0.167 0.067 0.039 0.037 0.007 0.007 0.003 0.001 0.001 0.001 0.897 0.861 Total Proportion of all cover 0.007 0.007 0.005 0.019 0.018 Bryophytes Eurynchiastrum pulchellum Brachythecium sp. Brachythecium sp. 1 Brachythecium sp. Pohlia nutans Dicranum sp. Funaria sp. Hylocomium splendens Marchantia sp. "Bottlebrush moss" Pleurozium schreberi Lichens Cladonia sp. Nephroma sp. Peltigera sp. 245 Cecil Lake Undisturbed Relevés - Total mean vegetation cover = 87.022% Species Trees Picea glauca Populus tremuloides Betula papyrifera Me an cover (%) Total Proportion of all cover 22.893 9.107 3.624 35.624 40.937 Total Proportion of all cover 6.908 6.762 4.388 4.015 1.922 0.971 0.782 0.445 0.426 0.297 0.279 0.190 0.115 0.035 0.029 0.003 27.567 31.678 Shrubs Rosa acicularis Viburnum edule Shepherdia canadensis Alnus viridis ssp. sinuata Cornus stolonifera Amelanchier alnifolia Rubus pubescens Ribes triste Symphoricarpos albus Ribes lacustre Lonicera dioica Salix sp. Lonicera involucrata Ribes oxyacanthoides Rubus idaeus Prunus virginiana Forbs Aralia nudicaulis Linnaea borealis Aster conspicuus Cornus canadensis Mertensia paniculata Lathyrus ochroleucus Clematis sp. Mitella nuda Disporum trachycarpum (aka Prosartes trachycarpa ) 246 6.460 3.137 1.118 1.058 0.349 0.146 0.133 0.108 0.086 Cecil Lake Undisturbed Relevés - cont'd Spe cie s Forbs - cont'd Orthilia secunda Geocaulon lividum Aster ciliolatus Galium boreale Pyrola chlorantha Pyrola asarifolia Fragaria virginiana Fragaria vesca Epilobium angustifolium Actaea rubra Osmorhiza chilensis Galium triflorum Viola sp. Vicia americana Delphinium glaucum Goodyera oblongifolia Listera s p. (aka Neottia sp.) Maianthemum canadense Taraxacum officinale Mean cover (%) Total Proportion of all cover 0.081 0.060 0.053 0.053 0.047 0.038 0.033 0.027 0.020 0.018 0.018 0.012 0.007 0.005 0.004 0.003 0.003 0.003 0.003 13.083 15.034 Total Proportion of all cover 2.253 0.253 0.240 2.746 3.156 Total Proportion of all cover 0.020 0.003 0.002 0.025 0.029 Graminoids "Fuzzy grass" "Thin grass" Calamagrostis canadensis Ferns & Fern Allies Fern sp. Equisetum arvense Gymnocarpium dryopteris 247 Cecil Lake Undisturbed Relevés - cont'd Species Bryophyte s Hylocomium splendens Pleurozium schreberi Ptilium crista-castrensis Rhytidiadelphus triquetrus Eurynchiastrum pulcellum Brachythecium sp. Moss sp. Dicranum polysetum Dicranum scoparium Plagiomnium cuspidatum Me an cover (%) Total Proportion of all cover 6.210 1.118 0.323 0.110 0.087 0.027 0.020 0.014 0.010 0.007 7.926 9.108 Total Proportion of all cover 0.047 0.003 0.050 0.057 Lichens Peltigera polydactylon Cladonia sp. 248 Hasler Flats Landslide Relevés - Total mean vegetation cover = 81.687% Species Mean cover (%) Trees Populus tremuloides 10.758 Populus balsamifera ssp. balsamifera 0.300 Betula papyrifera 0.103 Picea glauca 0.063 Total 11.224 Proportion of all cover 13.740 Shrubs Rosa acicularis 10.375 Alnus viridis ssp. sinuata 6.308 Rubus idaeus 5.595 Cornus stolonifera 4.900 Symphoricarpos albus 4.145 Viburnum edule 2.423 Lonicera dioica 2.348 Amelanchier alnifolia 1.528 Prunus virginiana 1.320 Lonicera involucrata 0.716 Ribes oxyacanthoides 0.461 Salix sp. 0.434 Rubus pubescens 0.348 Ribes triste 0.335 Salix sp. 0.266 Rubus parviflorus 0.019 Shepherdia canadensis 0.003 Ribes hudsonianum 0.001 Ribes laxiflorum 0.001 Total 41.526 Proportion of all cover 50.836 Forbs Smilacina racemosa 1.623 Aralia nudicaulis 1.403 Aster conspicuus 0.925 Mertensia paniculata 0.281 Petasites frigidus 0.248 249 Hasler Flats Landslide Relevés - cont'd Species Forbs - cont'd Vicia americana Streptopus amplexifolius Viola sp. Aster ciliolatus Lathyrus ochroleucus Galium boreale Heracleum lanatum Actaea rubra Smilacina stellata Taraxacum officinale Fragaria vesca Epilobium angustifolium Cornus canadensis Mitella nuda Galium triflorum Pyrola asarifolia Thalictrum occidentale Fragaria virginiana Cirsium arvense Spiraea betulifolia Maianthemum canadense Sonchus arvensis Epilobium ciliatum Linnaea borealis Veratrum viride Typha sp. Veronica beccabunga Cirsium vulgare Achillea millefolium Cyclamen hederifolium Hieracium sabaudum Orthilia secunda Geum macrophyllum Chenopodium capitatum Mean cover (%) 0.247 0.232 0.199 0.176 0.176 0.163 0.098 0.090 0.088 0.088 0.084 0.082 0.074 0.068 0.065 0.061 0.054 0.047 0.046 0.040 0.039 0.030 0.029 0.027 0.023 0.015 0.010 0.007 0.005 0.003 0.003 0.003 0.002 0.001 250 Hasler Flats Landslide Relevés - cont'd Species Forbs - cont'd Boechera stricta Plantago major "Pondweed" Trifolium hybridum Total Proportion of all cover Graminoids Calamagrostis canadensis Bromus inermis Elymus glaucus ssp. glaucus Poa palustris "Oat grass" Bromus ciliatus Leymus innovatus Elymus trachycaulus ssp. trachycaulus Carex sp. Carex rostrata Festuca sp. Trisetum cernuum "Limp grass" "Frothy grass" Phleum pratense Total Proportion of all cover Ferns & Fern allies Equisetum arvense Equisetum variegatum Fern sp. Total Proportion of all cover Mean cover (%) 0.001 0.001 0.001 0.001 6.859 8.397 1.088 1.051 0.696 0.038 0.019 0.016 0.012 0.010 0.010 0.008 0.007 0.005 0.005 0.003 0.001 2.969 3.635 16.417 0.002 0.002 16.421 20.102 251 Hasler Flats Landslide Relevés - cont'd Species Bryophytes Pohlia nutans Marchantia polymorpha Dicranum polysetum Syntrichia ruralis Plagiomnium cuspidatum Leptobryum pyriforme Total Proportion of all cover Mean cover (%) 2.423 0.198 0.062 0.004 0.004 0.002 2.693 3.297 252 Hasler Flats Undisturbed Relevés - Total mean vegetation cover = 100.807% Species Trees Populus tremuloides Populus balsamifera ssp. balsamifera Picea glauca Acer glabrum Mean cover (%) Total Proportion of all cover 13.580 4.335 0.510 0.033 18.458 18.310 Total Proportion of all cover 9.863 7.358 6.983 5.378 5.283 2.503 1.976 1.609 1.236 1.026 0.904 0.551 0.487 0.438 0.195 0.130 0.083 0.033 0.003 0.001 46.040 45.671 Shrubs Alnus viridis ssp. sinuata Cornus stolonifera Viburnum edule Symphoricarpos albus Rosa acicularis Amelanchier alnifolia Lonicera dioica Prunus virginiana Rubus idaeus Salix sp. 2 Ribes oxyacanthoides Rubus pubescens Ribes triste Lonicera involucrata Salix sp. 1 Ribes hudsonianum Shepherdia canadensis Salix sp. 3 Rubus parviflorus Ribes lacustre Forbs Aralia nudicaulis Heracleum lanatum Aster conspicuus Mitella nuda 6.856 3.844 2.556 1.397 253 Hasler Flats Undisturbed Relevés - cont'd Species Forbs - cont'd Smilacina racemosa Actaea rubra Thalictrum occidentale Smilacina stellata Viola sp. Lathyrus ochroleucus Maianthemum canadense Streptopus amplexifolius Vicia americana Petasites frigidus Angelica lucida Pyrola asarifolia Aster foliaceus Mertensia paniculata Cirsium arvense Aster ciliolatus Galium boreale Galium triflorum Disporum trachycarpum (aka Prosartes trachycarpa ) Fragaria vesca Osmorhiza chilensis Cornus canadensis Orthilia secunda Fragaria virginiana Taraxacum officinale Epilobium ciliatum Veronica beccabunga Veratrum viride Linnaea borealis Spiraea betulifolia Epilobium angustifolium Geum macrophyllum 254 Mean cover (%) 0.653 0.580 0.516 0.488 0.415 0.328 0.327 0.276 0.235 0.231 0.199 0.163 0.131 0.124 0.123 0.110 0.085 0.077 0.074 0.056 0.035 0.034 0.032 0.027 0.026 0.019 0.018 0.014 0.011 0.005 0.003 0.003 Hasler Flats Undisturbed Relevés - cont'd Species Forbs - cont'd Cyclamen hederifolium Achillea millefolium Monotropa uniflora Mean cover (%) Total Proportion of all cover Graminoids Calamagrostis canadensis "Mystery grass 2" Bromus inermis ssp. inermis "Hairy grass" Carex disperma "Mystery grass" Bromus ciliatus Poa pratensis Leymus innovatus Elymus lanceolatus ssp. lanceolatus Phleum pratense "Soft grass" Unknown grass sp. Carex sp. Glyceria sp. 0.002 0.001 0.001 20.075 19.914 Total Proportion of all cover 0.185 0.077 0.028 0.024 0.018 0.017 0.008 0.008 0.005 0.005 0.003 0.003 0.003 0.003 0.002 0.389 0.386 Total Proportion of all cover 15.625 0.007 0.002 15.634 15.509 Total Proportion of all cover 0.105 0.068 0.041 0.214 0.212 Ferns & Fern allies Equisetum arvense Equisetum pratense Botrychium virginianum Bryophytes Dicranum polysetum Plagiomnium cuspidatum Plagiomnium ellipticum 255 Appendix 4 Landslide geotypes glossary The geotype concept was developed in this study to delineate and classify geomorphic diversity as it relates to potential microsite and ecosystem diversity. No explicit guidelines existed for this work; therefore, a novel classification system was designed, using various sources for reference. The geotype definitions draw from several disciplines, including geography, geology, geomorphology, and hydrology. The main feature of the geotype classifications is that they describe identification of features on digital LiDAR imagery (rather than in the field), with the assistance of a digital elevation model. AP – Apron A relatively gentle, fanned out slope at the foot of a steeper slope, which is formed by materials from the steeper, upper slope. BA – Bank The sides of the channel of a river or creek, as well as the level land adjacent. BE – Bench A long, relatively narrow strip of level or gently sloped land with distinctly steeper slopes above and below it. BL – Blocky site An accumulation of large (>25 cm) chunks of consolidated, mostly unvegetated substrate. BS – Sandstone bedrock Exposed sedimentary rock, identified as sandstone based on background knowledge of local stratigraphy. CU – Cultivated field Raft of material broken off and transported from adjacent agricultural field. DF – Debris flow A mass of loose mud, sand, soil, rocks, vegetation, and air progressing down a slope. Seepage is present, and a water source is evident. DP – Depression A low spot, identified by a darker circle compared to surrounding terrain. 256 EF – Earthflow A moving mass of saturated fine-grained materials (eg. clay, fine sand, silt) progressing downslope. Visually distinguished from debris flows by the lack of large materials in the matrix. FL – Fluvial A river or creek channel with flowing water evident. Fl – Flood deposits – seasonal Fine blanket of material deposited by high water events. Identified by paler colour and flat appearance compared to surrounding terrain. FP – Floodplain A relatively flat, extensive area next to a river or creek, where water seasonally rises and covers the terrain. Identified by pools of standing water and shorter, saturation-tolerant vegetation. GU – Gully Sharp erosions of soil due to running water, usually on a hill side. Can be tens of metres wide and deep. HU – Hummocks Clusters of small knolls or mounds, usually <15 m tall. PI – Pillar A tall vertical column of slide material that remains after surrounding material has wasted away. PL – Plateau A flat, elevated landform rising distinctly above the surrounding terrain on at least one side. PO – Pond/wet area Any depression that contains standing water for any period of time, as visible on drone imagery. There is no lower size or depth limit. Identified in imagery by much darker colour than surrounding terrain and/or presence of cattail (Typha spp.). RA – Raft Intact pieces of vegetated terrain that have been transported down the landslide. Can consist of surrounding mature forest, grassland, or cultivated field. RB – Rotational blocks Blocks of slide material that are tilted due to rotational movement. 257 RF – Rockfall/topple A concentrated accumulation of rock that has fallen freely from a cliff face and moved downslope. RI – Ridge A chain of eroded hills that form a continuous elevated crest for some distance. RU – Rubble A blanket of irregular broken stone fragments on gentle or flat terrain. SB – Sandbar A ridge or island of sand in a river or creek channel. SC – Scarp A steep, often sparsely vegetated surface at the upper edge or within the landslide, caused by movement of displaced material away from stable material. SG – Steep grassy slope A stable slope generally > 35% gradient, covered in grasses and some short shrubs. SM – Slide matrix The main portion of the slide material within which other geomorphic features are found. Usually sparsely vegetated regosol. ST – Steep treed slope A stable slope generally > 35% gradient, covered in mature trees. SW – Swale/wrinkle Gently rolling lateral extensions of material populated by thick shrub growth. TA – Talus/scree A sheet of loose rock fragments over a slope. 258 Appendix 5 Geotype polygons Beatton River Landslide Geotype polygons Geotype code Description AP Apron - colluvial BL Blocky BL Blocky BL Blocky BL Blocky CU Cultivated field CU Cultivated field DF Debris flow FL Fluvial deposit GU Gully HU Hummocks PO Pond/wet area PO Pond/wet area PO Pond/wet area PO Pond/wet area PO Pond/wet area PO Pond/wet area PO Pond/wet area PO Pond/wet area PO Pond/wet area PO Pond/wet area PO Pond/wet area PO Pond/wet area PO Pond/wet area PO Pond/wet area PO Pond/wet area RA Raft RA Raft RA Raft RA Raft RA Raft RA Raft RA Raft RA Raft Area (m2 ) Index in Layer 9240 39 1447 26 4889 0 5550 22 7630 27 27.9 15 197.3 12 2996 11 165.6 18 370.7 36 43060 34 8 54 8.8 46 9.8 52 9.9 47 13.4 48 21.9 49 32.4 53 35.7 43 60.4 51 72.4 42 77.2 50 91.6 41 114.4 45 129.9 44 132.3 40 67.7 6 197.9 30 518 19 610 32 670 16 718 14 938 29 1492 2 259 Beatton River Landslide Geotype polygons - continued RA RA RA RA RA RB RB RF RF RF RF RU RU RU SB SC SC SC SC SC SC Raft Raft Raft Raft Raft Rotational block Rotational block Rockfall/topple Rockfall/topple Rockfall/topple Rockfall/topple Rubble Rubble Rubble Sandstone bedrock Scarp Scarp Scarp Scarp Scarp Scarp 2110 2899 3014 3560 9010 211.6 844 859 2335 3102 4519 364.8 7440 11890 802 439.1 591 2312 3011 5360 15960 13 17 28 23 7 9 1 4 3 31 5 35 37 21 10 24 33 8 20 38 25 Beatton River Undisturbed Geotype polygons Ge otype code CU DP FP PO PO PO RU SB SG ST ST Area (m2) Index in Layer Description Cultivated field 11750 0 Depression 9250 3 Floodplain 65000 10 Pond/wet area 606 6 Pond/wet area 340.5 7 Pond/wet area 97.5 8 Rubble 11900 2 Sandbar 9310 1 Steep grassy slope 99100 4 Steep treed slope 30250 5 Steep treed slope 64600 9 260 Cecil Lake Landslide Geotype polygons Geotype code EF FL FL FL Fl Fl Fl Fl GU HU HU PI PI PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO Area (m2 ) Index in Layer Description Earth flow 240 13 Fluvial 500 312 Fluvial 1135 319 Fluvial 4354 311 Flood deposits -seasonal 543 315 Flood deposits -seasonal 868 314 Flood deposits -seasonal 1108 313 Flood deposits -seasonal 4052 19 Gully 222.8 12 Hummocks 2520 0 Hummocks 7790 1 Pillar 135.2 17 Pillar 143.9 18 Pond/wet area 0.7 55 Pond/wet area 0.7 222 Pond/wet area 0.8 58 Pond/wet area 0.8 200 Pond/wet area 0.8 219 Pond/wet area 0.9 50 Pond/wet area 0.9 57 Pond/wet area 0.9 210 Pond/wet area 1 94 Pond/wet area 1 180 Pond/wet area 1 192 Pond/wet area 1 198 Pond/wet area 1 216 Pond/wet area 1 224 Pond/wet area 1.1 166 Pond/wet area 1.1 217 Pond/wet area 1.3 201 Pond/wet area 1.4 177 Pond/wet area 1.6 56 Pond/wet area 1.6 197 Pond/wet area 1.8 129 261 Cecil Lake Landslide Geotype polygons - continued Geotype code PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO Description Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Area (m2 ) Index in Layer 1.8 227 1.9 43 2 178 2.1 148 2.1 175 2.2 54 2.2 236 2.3 52 2.4 157 2.5 37 2.5 220 2.5 225 2.7 51 2.7 114 2.7 174 2.7 179 2.7 223 2.8 61 2.8 221 2.9 205 3.1 199 3.2 228 3.3 38 3.4 49 3.4 102 3.5 30 3.5 122 3.6 209 3.7 176 3.9 86 3.9 111 3.9 133 3.9 230 4 35 262 Cecil Lake Landslide Geotype polygons - continued Geotype code PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO Description Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Area (m2 ) Index in Layer 4 208 4.1 118 4.3 202 4.4 110 4.4 207 4.5 146 4.5 161 4.6 135 4.6 215 4.6 218 4.7 112 4.7 116 4.7 181 4.8 66 4.8 137 4.9 143 5 41 5.1 65 5.1 206 5.2 36 5.2 132 5.3 79 5.5 282 5.6 144 5.6 167 5.6 170 5.7 93 5.9 60 6 46 6 117 6 125 6.1 140 6.2 194 6.3 78 263 Cecil Lake Landslide Geotype polygons - continued Geotype code PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO Description Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Area (m2 ) Index in Layer 6.3 81 6.3 171 6.5 168 6.5 226 6.6 203 6.6 212 6.7 238 6.8 136 6.9 47 7.1 34 7.1 204 7.2 237 7.7 113 7.9 53 8 97 8.1 59 8.2 45 8.3 300 8.5 40 8.5 100 8.6 274 8.7 147 8.8 101 8.9 134 9 44 9.1 138 9.7 156 10.1 301 10.4 164 10.6 191 10.9 235 11 119 11.2 42 11.4 92 264 Cecil Lake Landslide Geotype polygons - continued Geotype code PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO Description Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Area (m2 ) Index in Layer 11.4 95 11.5 303 11.6 173 11.8 32 12 169 12.2 77 12.3 149 12.4 145 12.5 29 13 162 13.5 232 13.8 39 13.9 172 14 70 14.6 121 14.7 229 15.1 165 15.2 245 15.5 275 16 128 16.3 67 16.8 76 16.8 98 16.8 182 17.2 297 17.2 48 17.2 163 17.4 160 17.5 63 17.8 159 17.9 142 17.9 270 18.3 83 18.5 73 265 Cecil Lake Landslide Geotype polygons - continued Geotype code PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO Description Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Area (m2 ) Index in Layer 18.9 273 19.3 33 20.2 247 20.4 62 20.9 272 21.3 103 21.5 75 22 213 22.1 261 22.7 260 22.8 120 23.6 151 24.5 107 24.6 195 24.8 126 25.4 87 25.6 211 26.4 187 27.4 233 28 105 28.7 280 29.1 302 29.1 234 29.3 306 29.4 82 30.4 256 30.7 186 31 27 31.1 296 31.2 71 31.2 269 32.1 64 32.1 255 32.4 271 266 Cecil Lake Landslide Geotype polygons - continued Geotype code PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO Description Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Area (m2 ) Index in Layer 32.7 72 32.8 295 33.4 239 34.5 246 34.9 74 34.9 281 35.5 307 35.7 141 35.7 183 36.1 189 36.7 289 38.8 299 40.4 304 40.5 254 41.5 80 41.9 264 42.2 158 44.4 104 45 109 45.6 90 46.4 190 46.7 196 46.8 139 47.3 88 47.3 294 47.6 130 48.2 106 49.2 279 51.8 193 52.1 89 57.5 131 58.4 309 61.3 276 64.2 286 267 Cecil Lake Landslide Geotype polygons - continued Geotype code PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO Description Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Area (m2 ) Index in Layer 66.5 278 67 115 67.2 85 67.4 127 68.4 99 68.6 108 69.8 310 71.8 288 75.9 259 77.9 184 79.2 305 82.6 152 88.2 284 91.3 185 92 231 97.1 285 98.8 84 100.8 96 102 287 109.8 124 110.1 253 114.3 68 114.3 123 115 28 117 91 118.8 188 126.3 250 128.9 290 129 248 129.2 263 138.2 298 138.7 214 142 244 147.9 150 268 Cecil Lake Landslide Geotype polygons - continued Geotype code PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO RA RA RA RA RA RA RA RA RA Description Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Raft Raft Raft Raft Raft Raft Raft Raft Raft Area (m2 ) Index in Layer 179.5 31 184.5 243 185.8 262 189.6 277 216.9 293 218.4 308 219.8 292 262.9 155 283.4 153 287.7 69 306.3 283 317.7 268 318.8 251 362.6 154 499.9 258 539 257 636 267 695 291 797 241 1115 252 1141 266 1424 249 2231 240 2281 242 5790 265 221.6 21 390.5 3 429 22 1009 2 1211 14 1277 5 1387 7 2827 322 4466 8 269 Cecil Lake Landslide Geotype polygons - continued Geotype code RA RA RI RI RI RI RI RI RI RI RI RI RI RI SC SC SW SW SW Description Raft Raft Ridge Ridge Ridge Ridge Ridge Ridge Ridge Ridge Ridge Ridge Ridge Ridge Scarp Scarp Swale/wrinkle Swale/wrinkle Swale/wrinkle Area (m2 ) Index in Layer 6840 6 17320 4 408.5 317 586 15 607 323 645 25 965 24 1667 324 2117 16 2244 23 2719 316 2786 20 5330 26 5790 318 2218 321 9490 320 617 10 902 11 1404 9 270 Cecil Lake Undisturbed Geotype polygons Geotype code BA CU CU FL FL FL FP PL PL PL PO PO PO PO SG ST ST TA TA TA 2 Area (m ) Index in Layer Description Bank 19210 17 Cultivated field 10290 0 Cultivated field 16380 1 Fluvial 177.8 7 Fluvial 314.1 4 Fluvial 4108 2 Floodplain 1014 19 Plateau 29450 12 Plateau 32980 14 Plateau 50700 16 Pond/wet area 28.1 11 Pond/wet area 437.9 10 Pond/wet area 570 8 Pond/wet area 842 9 Steep grassy slope 184300 15 Steep treed slope 13660 13 Steep treed slope 144200 18 Talus/scree 957 6 Talus/scree 17440 3 Talus/scree 35140 5 271 Hasler Flats Landslide Geotype polygons Ge otype code PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO PO RA RA RA RA RA RA RA RA RI RI SC SC Description Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Pond/wet area Raft Raft Raft Raft Raft Raft Raft Raft Ridge Ridge Scarp Scarp Area (m2) Index in Layer 4.6 5.2 6.3 7.2 7.6 9.4 10.3 10.4 10.8 12.4 15.1 17.5 22.6 26 38 38.1 39.9 53.3 55.5 58.2 87.6 104.3 105 111.6 134.5 490.3 121 132.9 287.6 366 383.4 486.1 616 4456 88.5 142 173.5 730 36 17 24 33 12 34 23 16 30 28 11 13 22 15 21 26 27 14 25 35 29 31 32 19 18 20 0 2 1 5 4 6 3 38 10 8 37 9 272 Hasler Flats Undisturbed Geotype polygons Ge otype code BE BE BE BE DP DP FP FP PL PO PO SC SC SC SC SC Area (m2) Index in Layer De scription Bench 440.2 4 Bench 385.4 9 Bench 335.5 11 Bench 750 14 Depression 428.8 7 Depression 497.9 13 Floodplain 1411 2 Floodplain 1760 3 Plateau 6190 5 Pond/wet area 16 0 Pond/wet area 7.5 1 Scarp 340.3 6 Scarp 661 8 Scarp 363.2 10 Scarp 352.1 12 Scarp 1046 15 273 Appendix 6 Microtopography elevation summary tables - BEC plots Beatton River BEC plot elevation summary (n = 28 plots) Plot BE1 BE2 BE3 BE4 BE5 BE6 BE7 BE8 BE9 BE10 BE11 BE12 BE13 BE14 BE15 BE16 BE17 BE18 BE19 BE20 BE21 BE22 BE23 BE24 BE25 BE26 BE27 BE28 No. of Mean points elev (m) 49 644.81 50 648.03 49 640.05 49 645.42 49 643.06 50 644.54 50 642.77 51 629.00 49 594.54 47 591.38 48 594.38 45 591.42 48 575.60 47 558.80 48 577.12 47 513.62 49 511.33 50 489.28 49 491.24 48 480.94 46 475.02 50 473.63 48 484.04 47 546.41 49 644.69 51 649.54 50 645.65 49 640.08 Mean Min Max Min Standardised Standardised microsite microsite microsite elev SD CV elev (m) elev (m) elev (m) (m) (+/-m) (%) 643.56 1.25 0 4.88 0.87 69.41 645.57 2.46 0 4.34 1.21 49.38 639.29 0.76 0 1.39 0.38 50.09 642.10 3.33 0 7.97 1.86 55.90 640.92 2.14 0 6.17 1.14 53.38 638.61 5.93 0 11.30 3.33 56.20 641.81 0.95 0 4.47 0.84 88.29 627.26 1.75 0 3.72 1.04 59.34 593.78 0.77 0 2.33 0.54 70.87 591.05 0.33 0 1.88 0.46 139.42 593.30 1.08 0 2.21 0.60 55.69 590.68 0.74 0 1.37 0.36 49.12 575.01 0.58 0 1.81 0.47 80.36 556.37 2.43 0 5.02 1.34 55.20 574.13 2.99 0 6.14 1.66 55.73 512.43 1.19 0 2.09 0.55 46.17 510.04 1.28 0 4.28 0.92 71.57 488.87 0.42 0 1.09 0.28 67.53 490.39 0.85 0 2.33 0.51 60.20 480.24 0.70 0 1.33 0.32 45.71 474.59 0.43 0 0.91 0.21 48.15 473.31 0.32 0 1.02 0.21 64.33 482.30 1.74 0 3.03 0.73 41.82 545.57 0.84 0 1.56 0.39 45.74 643.36 1.34 0 2.90 0.92 69.12 645.96 3.58 0 7.25 2.04 56.98 645.17 0.48 0 0.99 0.24 49.85 636.22 3.86 0 7.58 2.02 52.19 274 Cecil Lake BEC plot elevation summary (n = 55 plots) Plot CE1 CE2 CE3 CE4 CE5 CE6 CE7 CE8 CE9 CE10 CE11 CE12 CE13 CE14 CE15 CE16 CE17 CE18 CE19 CE20 CE21 CE22 CE23 CE24 CE25 CE26 CE27 CE28 CE29 CE30 No. of Mean points elev (m) 49 670.75 46 661.89 48 646.31 48 647.64 46 647.60 49 633.92 49 629.59 50 663.58 48 658.92 48 663.48 46 658.25 48 647.26 49 655.66 48 646.35 51 641.23 50 642.02 46 632.64 49 629.66 50 626.75 51 614.60 47 621.29 49 652.20 50 644.48 46 638.38 49 636.20 50 634.52 51 627.78 48 616.41 48 600.86 46 579.89 Mean Min Max Min Standardised Standardised elev microsite microsite microsite SD CV elev (m) elev (m) elev (m) (m) (+/-m) (%) 669.25 1.50 0 3.54 0.93 62.48 660.42 1.47 0 2.44 0.68 46.45 646.08 0.23 0 0.84 0.19 79.53 646.23 1.41 0 2.91 0.89 63.39 645.41 2.19 0 3.63 1.11 50.66 631.44 2.48 0 4.77 1.38 55.63 629.17 0.42 0 0.78 0.22 52.74 661.83 1.75 0 3.04 0.83 47.07 657.26 1.66 0 2.66 0.73 44.05 662.53 0.95 0 1.89 0.50 53.23 656.34 1.91 0 3.76 1.17 61.18 645.07 2.19 0 4.71 1.27 58.05 653.88 1.78 0 2.76 0.78 43.95 645.84 0.51 0 0.73 0.15 29.17 640.83 0.40 0 0.72 0.18 46.01 641.23 0.79 0 1.40 0.29 37.31 631.10 1.54 0 3.39 1.01 65.54 628.45 1.21 0 1.80 0.47 38.93 626.00 0.75 0 1.25 0.28 37.60 614.45 0.15 0 0.54 0.12 74.66 620.42 0.87 0 1.80 0.45 51.40 650.59 1.61 0 2.59 0.74 45.71 643.92 0.56 0 0.88 0.23 41.73 637.38 1.00 0 1.88 0.54 54.41 635.57 0.63 0 1.42 0.37 59.57 634.35 0.17 0 0.87 0.19 112.80 627.42 0.36 0 1.03 0.25 69.62 615.01 1.40 0 2.60 0.76 54.15 600.26 0.60 0 1.64 0.39 65.89 578.68 1.21 0 2.23 0.57 46.81 275 Cecil Lake BEC plot elevation summary - cont'd Plot CE31 CE32 CE33 CE34 CE36 CE37 CE38 CE39 CE40 CE41 CE42 CE43 CE44 CE45 CE46 CE47 CE48 CE49 CE50 CE51 CE52 CE53 CE54 CE55 CE56 No. of Mean points elev (m) 50 564.17 50 568.46 50 578.46 48 574.87 50 640.36 48 645.77 49 646.10 50 641.34 48 639.52 52 626.70 49 611.79 50 610.97 50 626.78 47 588.08 52 633.83 48 632.16 48 616.63 47 588.08 51 580.27 49 578.59 49 572.10 48 572.15 49 570.99 49 582.11 49 581.21 Mean Min Max Min Standardised Standardised elev microsite microsite microsite SD CV elev (m) elev (m) elev (m) (m) (+/-m) (%) 564.08 0.09 0 0.21 0.04 43.51 567.72 0.74 0 1.36 0.38 52.25 577.21 1.25 0 2.55 0.79 63.03 574.77 0.10 0 0.14 0.02 22.46 640.28 0.08 0 0.15 0.03 40.87 644.88 0.89 0 1.88 0.56 62.80 644.99 1.11 0 1.93 0.54 49.08 640.08 1.26 0 2.19 0.57 45.74 638.94 0.58 0 0.88 0.18 30.82 626.57 0.13 0 0.47 0.09 72.30 611.35 0.44 0 1.10 0.32 72.94 610.76 0.21 0 1.23 0.25 118.97 625.46 1.32 0 2.60 0.84 63.42 586.01 2.07 0 4.05 1.08 52.10 633.03 0.80 0 1.45 0.37 45.97 631.60 0.56 0 1.47 0.43 77.63 616.02 0.61 0 1.53 0.36 59.16 586.01 2.07 0 4.05 1.08 52.10 579.31 0.96 0 1.96 0.62 65.05 575.91 2.68 0 4.32 1.24 46.13 571.97 0.13 0 0.54 0.10 72.33 571.18 0.97 0 1.48 0.34 34.63 570.47 0.52 0 1.10 0.29 55.65 581.57 0.54 0 1.14 0.30 55.37 580.97 0.24 0 0.91 0.21 88.92 276 Hasler Flats BEC plot elevation summary (n = 29) No. of Mean points elev Plot (m) HA2 41 598.72 HA3 51 599.56 HA4 52 601.95 HA5 62 599.19 HA6 65 601.10 HA7 99 609.13 HA8 74 608.47 HA9 66 606.40 HA10 43 606.87 HA11 48 604.89 HA12 36 603.79 HA13 58 604.59 HA14 74 603.80 HA15 90 604.29 HA16 102 601.01 HA17 79 601.15 HA18 92 601.81 HA19 70 599.28 HA20 69 600.72 HA21 23 600.69 HA22 36 598.98 HA23 29 597.25 HA24 60 599.87 HA25 52 600.39 HA26 42 602.12 HA27 81 603.00 HA28 58 607.83 HA29 112 602.58 HA30 78 601.15 Mean Min Max Standardised Standardised Min SD CV elev microsite microsite microsite elev (m) elev (m) elev (m) (+/-m) (%) (m) 598.24 0.48 0 1.24 0.26 54.05 598.74 0.82 0 1.74 0.47 57.18 601.33 0.62 0 1.51 0.40 64.81 598.17 1.01 0 2.55 0.73 71.48 600.41 0.69 0 1.38 0.29 41.77 607.97 1.16 0 2.82 0.78 67.24 606.90 1.57 0 2.48 0.68 43.32 605.33 1.07 0 1.93 0.56 52.16 606.50 0.37 0 0.82 0.18 48.51 604.59 0.30 0 0.55 0.13 42.29 603.38 0.41 0 1.81 0.38 91.17 603.77 0.82 0 1.86 0.50 60.75 603.17 0.63 0 1.03 0.21 34.09 602.92 1.37 0 3.18 0.84 61.53 600.00 1.01 0 1.64 0.46 45.51 599.94 1.21 0 1.81 0.53 44.20 600.75 1.06 0 1.57 0.34 31.77 598.76 0.52 0 1.09 0.25 48.00 600.20 0.52 0 1.09 0.28 53.71 599.93 0.76 0 1.04 0.23 29.93 598.74 0.24 0 0.67 0.16 64.27 597.13 0.12 0 0.22 0.06 51.50 599.09 0.78 0 1.43 0.33 41.90 599.83 0.56 0 1.05 0.33 60.08 600.58 1.54 0 2.34 0.54 35.14 602.00 1.00 0 1.96 0.54 53.84 606.65 1.18 0 4.23 1.16 98.30 601.63 0.95 0 1.90 0.59 62.31 600.87 0.28 0 0.66 0.17 61.94 277 Pond Pond Slide Size Slide UTMs Slide Type # # (ha) 0.0009 515 141 10U 635411 6231647 Retrogressive rotational failure 0.002 154 61 10U 648617 6262829 Retrogressive rotational failure 0.0023 517 141 10U 635411 6231647 Retrogressive rotational failure 0.0026 680 196 10U 658130 6232188 Retrogressive rotational failure 0.0026 52 71 10U 660889 6229983 Shallow retrogressive failure 0.0028 96 45 10U 645103 6237317 Retrogressive rotational failure 0.003 51 71 10U 660889 6229983 Shallow retrogressive failure 0.0032 616 173 10U 647595 6263055 Compound failure 0.0032 153 61 10U 648617 6262829 Retrogressive rotational failure 0.0034 497 136 10U 627698 6232647 Rotational failure 0.0034 587 47 10U 640425 6241038 Retrogressive rotational failure 0.0036 88 40 10U 648407 6238441 Multi-level rotational failure 0.0036 344 94 10U 586742 6231586 Rotational failure 0.004 280 83 10U 611833 6238026 Retrogressive rotational failure 0.0043 424 113 10U 607910 6245770 Rotational failure 0.0043 378 100 10U 598414 6248139 Compound failure 0.0045 48 71 10U 660889 6229983 Shallow retrogressive failure 0.0046 679 196 10U 658130 6232188 Retrogressive rotational failure 0.0047 283 83 10U 611833 6238026 Retrogressive rotational failure 0.0049 82 40 10U 648407 6238441 Multi-level rotational failure 0.005 37 27 10U 661452 6225535 Compound failure 0.005 367 100 10U 598414 6248139 Compound failure 0.0051 476 126 10U 631665 6307193 Rotational failure 0.0054 456 122 10U 619929 6209531 Mobile flow 0.0054 345 94 10U 586742 6231586 Rotational failure 0.0054 519 141 10U 635411 6231647 Retrogressive rotational failure 0.0055 520 141 10U 635411 6231647 Retrogressive rotational failure 0.0056 87 41 10U 647664 6238258 Retrogressive rotational failure 0.0056 522 140 10U 634447 6231204 Retrogressive rotational failure 0.0059 514 141 10U 635411 6231647 Retrogressive rotational failure 0.006 342 94 10U 586742 6231586 Rotational failure 0.006 521 141 10U 635411 6231647 Retrogressive rotational failure Pond location on slide Flat area midway down slide In fresh material Flat area midway down slide Right next to creek Just below headscarp (small, intact) Below small scarp (intact) Below headscarp (small, intact) Just below new part of headscarp (small, intact) In fresh material Above small scarp (intact), near river Above small scarp (in debris) Debris apron Debris apron Toe Below small scarp (in debris) Debris apron near river Just below headscarp (small, intact) Next to creek At bottom of slide Debris apron below large scarp (intact) Toe Debris apron near river Base of headscarp (small, intact) Near top of slide Debris apron Above small scarp (intact) Base of slide, next to creek Below small scarp (intact) Above small scarp (intact) Flat area midway down slide Debris apron Below small scarp (intact) 278 Dormant Active Dormant Dormant Dormant Dormant Dormant Active Active Dormant Dormant Active Dormant Dormant Dormant Dormant Dormant Dormant Dormant Active Dormant Dormant Dormant Active Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Slide Activity Appendix 7 Landslide pond descriptions - Mapsheet 94A Geomorphic location of pond Body Toe Toe Toe Head Toe Body Head Toe Body Toe Toe Body Toe Head Toe Head Toe Toe Toe Toe Toe Head Body Body Toe Toe Body Body Body Body Body Pond Pond Slide Size Slide UTMs Slide Type # # (ha) 0.006 678 196 10U 658130 6232188 Retrogressive rotational failure 0.0061 457 122 10U 619929 6209531 Mobile flow 0.0063 36 27 10U 661452 6225535 Compound failure 0.0064 90 43 10U 646755 6238196 Retrogressive rotational failure 0.0064 317 89 10U 583121 6249823 Multi-level rotational failure 0.0064 381 100 10U 598414 6248139 Compound failure 0.0066 100 46 10U 640572 6238860 Retrogressive rotational failure 0.0066 637 182 10U 653926 6268718 Retrogressive rotational failure 0.0067 459 122 10U 619929 6209531 Mobile flow 0.0068 506 139 10U 633271 6230890 Compound failure 0.0068 25 60 10U 647207 6252566 Rotational failure 0.0068 4 57 10U 645882 6252696 Retrogressive rotational failure 0.0068 50 71 10U 660889 6229983 Shallow retrogressive failure 0.0069 335 90 10U 584584 6239849 Retrogressive rotational failure 0.0069 767 226 10U 673115 6239339 Multi-level rotational failure 0.0069 393 104 10U 607196 6251797 Compound failure 0.007 49 71 10U 660889 6229983 Shallow retrogressive failure 0.0071 484 130 10U 633609 6300494 Compound failure 0.0072 379 100 10U 598414 6248139 Compound failure 0.0072 615 173 10U 647595 6263055 Compound failure 0.0073 592 162 10U 639763 6241926 Retrogressive rotational failure 0.0074 542 3 10U 635265 6222327 Retrogressive rotational failure 0.0075 77 44 10U 646758 6237400 Retrogressive rotational failure 0.0075 369 100 10U 598414 6248139 Compound failure 0.0077 38 27 10U 661452 6225535 Compound failure 0.0077 377 100 10U 598414 6248139 Compound failure 0.0079 639 184 10U 657390 6240045 Rotational failure 0.0079 584 161 10U 639518 6240605 Retrogressive rotational failure 0.0081 423 113 10U 607910 6245770 Rotational failure 0.0081 530 144 10U 626024 6221319 Rotational failure 0.0081 635 182 10U 653926 6268718 Retrogressive rotational failure 0.0081 410 108 10U 607734 6248896 Rotational failure Pond location on slide Next to creek Near top of slide Toe Just above small scarp (intact) Debris apron, right at base of large scarp (intact) Debris apron near river Below small scarp (in debris) Below/within small scarp (intact), near river Near top of slide (midway across) On hillside in small depression Debris apron Below small scarp (intact) Just below headscarp (small, intact) Just above small retrogressive scarp (intact) Above small scarp (intact) Debris apron Just below headscarp (small, intact) Base of headscarp (small, intact) Debris apron Just below new part of headscarp (small, intact) Below headscarp (large, intact) Below small scarp (in debris) Below small scarp (in debris) Debris apron near river Toe Debris apron near river Debris apron near creek Below small scarp (intact), near river Base of headscarp (large, intact) Below small scarp (intact) Above small scarp (intact), near river Above small scarp (intact) 279 Dormant Active Dormant Active Dormant Dormant Dormant Dormant Active Dormant Dormant Active Dormant Dormant Dormant Dormant Dormant Dormant Dormant Active Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Slide Activity Appendix 7 Landslide pond descriptions - Mapsheet 94A - cont'd Geomorphic location of pond Toe Body Toe Toe Toe Toe Body Toe Body Body Toe Toe Body Body Body Body Body Head Body Head Head Body Body Toe Toe Toe Toe Toe Head Body Toe Body Pond Pond Slide Size Slide UTMs Slide Type # # (ha) 0.0083 417 111 10U 608499 6247479 Compound failure 0.0083 660 189 10U 654871 6235759 Retrogressive rotational failure 0.0084 373 100 10U 598414 6248139 Compound failure 0.0086 737 15 10U 655647 6215120 Retrogressive rotational failure 0.0087 402 105 10U 607741 6250210 Compound failure 0.0087 16 57 10U 645882 6252696 Retrogressive rotational failure 0.0088 472 126 10U 631665 6307193 Rotational failure 0.0089 659 189 10U 654871 6235759 Retrogressive rotational failure 0.0089 341 94 10U 586742 6231586 Rotational failure 0.0089 430 114 10U 607383 6243249 Retrogressive rotational failure 0.009 421 112 10U 608153 6247351 Rotational failure 0.009 607 53 10U 641413 6249461 Retrogressive rotational failure 0.009 617 173 10U 647595 6263055 Compound failure 0.0091 498 136 10U 627698 6232647 Rotational failure 0.0093 316 89 10U 583121 6249823 Multi-level rotational failure 0.0093 586 47 10U 640425 6241038 Retrogressive rotational failure 0.0094 548 145 10U 634428 6225821 Multi-level rotational failure 0.0095 47 71 10U 660889 6229983 Shallow retrogressive failure 0.0097 231 4 10U 637998 6220826 Multi-level rotational failure 0.0097 730 14 10U 655144 6217752 Compound failure 0.0098 346 94 10U 586742 6231586 Rotational failure 0.01 57 33 10U 657471 6230119 Compound failure 0.01 408 107 10U 607530 6250024 Compound failure 0.01 609 169 10U 642693 6252730 Retrogressive rotational failure 0.01 766 225 10U 672724 6237484 Retrogressive rotational failure 0.0101 20 57 10U 645882 6252696 Retrogressive rotational failure 0.0101 471 126 10U 631665 6307193 Rotational failure 0.0102 588 47 10U 640425 6241038 Retrogressive rotational failure 0.0102 254 1 10U 631931 6221078 Retrogressive rotational failure 0.0103 418 111 10U 608499 6247479 Compound failure 0.0104 163 22 10U 668246 6222087 Compound failure 0.0106 286 83 10U 611833 6238026 Retrogressive rotational failure Pond location on slide Above small scarp (in debris) Above small scarp (intact) Debris apron near river Next to South side scarp Within small scarp (in debris) below headscarp (small, intact) Just below headscarp (large, intact) Below headscarp (small, intact) Above small scarp (intact) Debris apron Near creek/river Below small scarp (intact) Below headscarp (large, intact) Below new part of headscarp (small, intact) Above small scarp (intact), near river Debris apron below large scarp (intact) Next to small scarp (in debris) Debris apron Just below headscarp (small, intact) Debris apron - near creek Within small scarp (in debris) Debris apron Below small scarp (intact) Within headscarp (small, intact) by south sidescarp Below headscarp (small, intact) Next to small scarp (intact) Near small scarp (in debris) Below headscarp (small, intact) Below small scarp (in debris), near river Above backtilt Below headscarp (large, intact) Below small scarp (intact) At bottom of slide 280 Dormant Dormant Dormant Dormant Dormant Active Dormant Dormant Dormant Dormant Dormant Dormant Active Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Active Dormant Dormant Dormant Dormant Dormant Dormant Slide Activity Appendix 7 Landslide pond descriptions - Mapsheet 94A - cont'd Geomorphic location of pond Body Body Toe Body Body Head Head Body Body Toe Toe Body Head Body Toe Toe Toe Head Toe Head Body Body Head Head Body Body Body Toe Body Head Body Toe Pond Pond Slide Size Slide UTMs Slide Type # # (ha) 0.0106 602 165 10U 641712 6246986 Rotational failure 0.0106 558 148 10U 635615 6217619 Retrogressive rotational failure 0.0106 372 100 10U 598414 6248139 Compound failure 0.0106 31 24 10U 662143 6223550 Rotational failure 0.011 404 105 10U 607741 6250210 Compound failure 0.0111 668 38 10U 652244 6236095 Retrogressive rotational failure 0.0112 502 137 10U 628079 6232225 Multi-level rotational failure 0.0112 550 145 10U 634428 6225821 Multi-level rotational failure 0.0113 309 87 10U 580867 6250762 Compound failure 0.0114 399 105 10U 607741 6250210 Compound failure 0.0118 368 100 10U 598414 6248139 Compound failure 0.0118 677 196 10U 658130 6232188 Retrogressive rotational failure 0.0118 749 214 10U 664400 6210645 Multi-level rotational failure 0.0122 376 100 10U 598414 6248139 Compound failure 0.0122 308 87 10U 580867 6250762 Compound failure 0.0123 219 4 10U 637998 6220826 Multi-level rotational failure 0.0124 478 127 10U 632593 6304450 Rotational failure 0.0125 518 141 10U 635411 6231647 Retrogressive rotational failure 0.0125 422 112 10U 608153 6247351 Rotational failure 0.0127 429 114 10U 607383 6243249 Retrogressive rotational failure 0.0127 638 183 10U 652608 6268058 Retrogressive rotational failure 0.0128 400 105 10U 607741 6250210 Compound failure 0.0128 489 132 10U 633416 6298822 Rotational failure 0.0129 724 207 10U 655366 6218822 Retrogressive rotational failure 0.0129 289 83 10U 611833 6238026 Retrogressive rotational failure 0.0129 425 113 10U 607910 6245770 Rotational failure 0.0129 535 1 10U 631931 6221078 Retrogressive rotational failure 0.013 371 100 10U 598414 6248139 Compound failure 0.013 509 140 10U 634447 6231204 Retrogressive rotational failure 0.0134 403 105 10U 607741 6250210 Compound failure 0.0134 512 141 10U 635411 6231647 Retrogressive rotational failure 0.0134 582 161 10U 639518 6240605 Retrogressive rotational failure Pond location on slide Base of headscarp (large, intact) Above small scarp (intact) Debris apron right by river Above small scarp (in debris) Above small scarp (in debris) Next to river at bottom of slide Just above small scarp (intact), near river Debris apron Debris apron - below headscarp (small, intact), above large scarp (intact) Within small scarp (in debris) below headscarp (small, intact) Debris apron near river Below small scarp (in debris), near creek Below headscarp (small, intact) Debris apron Just below headscarp (small, intact), above large scarp (intact) Next to small scarp (intact) Base of headscarp (small, intact) Above small scarp (intact) Above small scarp (intact) Near creek/river Below/within small scarp (intact) Below small scarp (in debris) Below small scarp (in debris), near river Within large scarp (intact) At bottom of slide Below small scarp (in debris) Below small scarp (in debris) Debris apron near river Below small scarp (intact) Below small scarp (in debris) Flat area midway down slide Below small scarp (intact), near river 281 Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Slide Activity Appendix 7 Landslide pond descriptions - Mapsheet 94A - cont'd Geomorphic location of pond Body Toe Toe Body Body Toe Body Toe Body Body Toe Toe Head Body Body Body Head Toe Body Toe Toe Body Toe Toe Toe Head Body Toe Body Body Body Toe Pond Pond Slide Size Slide UTMs Slide Type # # (ha) 0.0135 99 46 10U 640572 6238860 Retrogressive rotational failure 0.0136 566 153 10U 637499 6223113 Rotational failure 0.0138 781 233 10U 682783 6224622 Rotational failure 0.014 723 207 10U 655366 6218822 Retrogressive rotational failure 0.0141 80 40 10U 648407 6238441 Multi-level rotational failure 0.0141 18 57 10U 645882 6252696 Retrogressive rotational failure 0.0142 470 125 10U 622324 6288717 Rotational failure 0.0144 44 71 10U 660889 6229983 Shallow retrogressive failure 0.0144 445 121 10U 608754 6224408 Rotational failure 0.0144 191 10 10U 652580 6220750 Retrogressive rotational failure 0.0144 419 111 10U 608499 6247479 Compound failure 0.0145 415 111 10U 608499 6247479 Compound failure 0.0145 681 196 10U 658130 6232188 Retrogressive rotational failure 0.0145 613 172 10U 641260 6254208 Rotational failure 0.0147 365 100 10U 598414 6248139 Compound failure 0.0147 416 111 10U 608499 6247479 Compound failure 0.0147 655 189 10U 654871 6235759 Retrogressive rotational failure 0.0148 511 141 10U 635411 6231647 Retrogressive rotational failure 0.0149 374 100 10U 598414 6248139 Compound failure 0.015 774 229 10U 674428 6241207 Retrogressive rotational failure 0.0151 527 143 10U 624881 6220596 Retrogressive rotational failure 0.0152 86 41 10U 647664 6238258 Retrogressive rotational failure 0.0154 294 84 10U 612699 6237705 Retrogressive rotational failure 0.0154 763 224 10U 673168 6235959 Retrogressive rotational failure 0.0155 630 180 10U 634582 6300315 Retrogressive rotational failure 0.0157 329 90 10U 584584 6239849 Retrogressive rotational failure 0.0157 732 14 10U 655144 6217752 Compound failure 0.0158 98 45 10U 645103 6237317 Retrogressive rotational failure 0.0159 631 180 10U 634582 6300315 Retrogressive rotational failure 0.0162 526 143 10U 624881 6220596 Retrogressive rotational failure 0.0163 204 14 10U 655144 6217752 Compound failure 0.0163 485 130 10U 633609 6300494 Compound failure Pond location on slide Just below small scarp (in debris) Within small scarp (in debris) Debris apron below headscarp (large, intact) Below small scarp (intact) Debris apron below large scarp (intact) Next to small scarp (in debris) Below headscarp (small, intact) Just below headscarp (small, intact) Debris apron near river Below small scarp (intact) Below headscarp (large, intact) Below small scarp (in debris) Below headscarp (large, intact) Debris apron below headscarp (large, intact) Debris apron near river Above small scarp (in debris) Beside creek Flat area below small scarp (in debris) Debris apron Within headscarp (small, intact) Below small scarp (intact) Below small scarp (intact) Below small scarp (in debris) Below small scarp (intact) Base of small scarp (intact) Just below small retrogressive scarp (intact) Below small scarp (in debris) Below small scarp (in debris) Below small scarp (intact) Below small scarp (intact) Below small scarp (in debris) Below small scarp (intact) 282 Dormant Active Dormant Dormant Active Active Dormant Dormant Dormant Dormant Dormant Dormant Dormant Active Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Slide Activity Appendix 7 Landslide pond descriptions - Mapsheet 94A - cont'd Geomorphic location of pond Body Body Toe Body Toe Body Body Head Toe Toe Head Toe Body Body Toe Body Toe Toe Body Head Toe Body Toe Body Body Body Body Body Body Toe Body Body Pond Pond Slide Size Slide UTMs Slide Type # # (ha) 0.0166 557 148 10U 635615 6217619 Retrogressive rotational failure 0.0168 420 111 10U 608499 6247479 Compound failure 0.0169 683 197 10U 659267 6231807 Rotational failure 0.0169 682 196 10U 658130 6232188 Retrogressive rotational failure 0.017 460 122 10U 619929 6209531 Mobile flow 0.017 363 99 10U 598103 6246815 Rotational failure 0.0173 414 110 10U 608193 6247991 Compound failure 0.0173 411 109 10U 608185 6248414 Compound failure 0.0173 667 38 10U 652244 6236095 Retrogressive rotational failure 0.0173 505 138 10U 632037 6230791 Rotational failure 0.0174 282 83 10U 611833 6238026 Retrogressive rotational failure 0.0174 664 191 10U 649843 6238652 Rotational failure 0.0174 326 90 10U 584584 6239849 Retrogressive rotational failure 0.0174 375 100 10U 598414 6248139 Compound failure 0.0175 562 151 10U 637222 6220172 Compound failure 0.0176 362 99 10U 598103 6246815 Rotational failure 0.0177 718 204 10U 654675 6219635 Retrogressive rotational failure 0.0181 742 212 10U 656103 6216700 Retrogressive rotational failure 0.0182 30 24 10U 662143 6223550 Rotational failure 0.0184 328 90 10U 584584 6239849 Retrogressive rotational failure 0.0187 382 100 10U 598414 6248139 Compound failure 0.0187 641 185 10U 656213 6239712 Rotational failure 0.0188 176 20 10U 679534 6219946 Rotational failure 0.0189 392 103 10U 589643 6282999 Rotational failure 0.019 474 126 10U 631665 6307193 Rotational failure 0.0191 531 144 10U 626024 6221319 Rotational failure 0.0191 263 83 10U 611833 6238026 Retrogressive rotational failure 0.0194 748 214 10U 664400 6210645 Multi-level rotational failure 0.0194 440 116 10U 606417 6242052 Rotational failure 0.0194 302 85 10U 612971 6237140 Retrogressive rotational failure 0.0195 315 89 10U 583121 6249823 Multi-level rotational failure 0.0196 469 125 10U 622324 6288717 Rotational failure Above small scarp (intact) Below small scarp (in debris) Debris apron at base of headscarp (large, intact) Just above small scarp (in debris) Near top of slide (midway across) Within small scarp (in debris) Below headscarp (small, intact) Debris apron near river/creek Next to river at bottom of slide Debris apron At bottom of slide Within headscarp (large, intact) Above small retrogressive scarp (intact) Debris apron right by river Middle of slide Within small scarp (in debris) Below headscarp (small, intact) Bottom of slide, near creek Above small scarp (intact) Within small retrogressive scarp (intact) Debris apron near river Base of headscarp (large, intact) Below headscarp (large, intact) Debris apron Base of headscarp (small, intact) Above small scarp (intact) Below headscarp (large, intact), near west sidescarp Above small scarp (in debris) Debris apron Below small scarp (in debris) Debris apron below large scarp (intact) Halfway down slide Dormant Dormant Dormant Dormant Active Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant 283 Pond location on slide Slide Activity Appendix 7 Landslide pond descriptions - Mapsheet 94A - cont'd Geomorphic location of pond Toe Toe Body Body Body Body Body Toe Toe Toe Toe Head Body Toe Body Body Head Toe Body Body Toe Body Body Body Head Body Body Body Toe Toe Toe Body Pond Pond Slide Size Slide UTMs Slide Type # # (ha) 0.0198 213 9 10U 639115 6218246 Rotational failure 0.02 634 182 10U 653926 6268718 Retrogressive rotational failure 0.0201 725 207 10U 655366 6218822 Retrogressive rotational failure 0.0201 481 127 10U 632593 6304450 Rotational failure 0.0204 686 198 10U 660187 6227739 Retrogressive rotational failure 0.0204 398 105 10U 607741 6250210 Compound failure 0.0204 327 90 10U 584584 6239849 Retrogressive rotational failure 0.0206 447 122 10U 619929 6209531 Mobile flow 0.0208 473 126 10U 631665 6307193 Rotational failure 0.0208 370 100 10U 598414 6248139 Compound failure 0.021 644 186 10U 655856 6238250 Retrogressive rotational failure 0.021 483 129 10U 633999 6301639 Rotational failure 0.0212 722 207 10U 655366 6218822 Retrogressive rotational failure 0.0212 250 1 10U 631931 6221078 Retrogressive rotational failure 0.0215 360 97 10U 597665 6245826 Rotational failure 0.0216 409 108 10U 607734 6248896 Rotational failure 0.0221 490 132 10U 633416 6298822 Rotational failure 0.0222 126 57 10U 645882 6252696 Retrogressive rotational failure 0.0223 23 57 10U 645882 6252696 Retrogressive rotational failure 0.0224 606 53 10U 641413 6249461 Retrogressive rotational failure 0.0224 338 93 10U 585645 6232574 Rotational failure 0.0225 593 74 10U 636436 6241818 Compound failure 0.0226 649 187 10U 655524 6236858 Multi-level rotational failure 0.0226 661 190 10U 653976 6235516 Retrogressive rotational failure 0.0226 278 83 10U 611833 6238026 Retrogressive rotational failure 0.0233 21 57 10U 645882 6252696 Retrogressive rotational failure 0.0235 500 136 10U 627698 6232647 Rotational failure 0.0235 687 199 10U 659140 6227440 Multi-level rotational failure 0.0237 380 100 10U 598414 6248139 Compound failure 0.024 395 105 10U 607741 6250210 Compound failure 0.024 84 41 10U 647664 6238258 Retrogressive rotational failure 0.0241 160 63 10U 656420 6272207 Rotational failure Pond location on slide Below small scarp (intact) Above small scarp (intact), near river Within large scarp (intact) Below headscarp (small, intact) Below small scarp (intact), right by river At base of headscarp (large, intact) Above small retrogressive scarp (intact) Near base of slide Below headscarp (small, intact) Debris apron near river Below large headscarp (intact) Just below headscarp (small) and above small scarp (both intact) Base of headscarp (small, intact) Above backtilt On level portion of headscarp (large, intact) Next to small scarp (intact) Below small scarp (in debris), near river Below small scarp (in debris) Below small scarp (in debris) Below headscarp (large, intact) Bottom of slide, by river Below headscarp (large, intact) Halfway down slide, by south sidescarp Apron at top of slide below headscarp (large, intact) Toe Below headscarp (large, intact) Above small scarp (intact), near river Below small scarp (in debris) Debris apron Base of headscarp (small, intact) Below small scarp (intact) Just below small scarp (in debris) 284 Dormant Dormant Dormant Dormant Dormant Dormant Dormant Active Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Active Active Dormant Active Dormant Dormant Dormant Dormant Active Dormant Dormant Dormant Dormant Dormant Dormant Slide Activity Appendix 7 Landslide pond descriptions - Mapsheet 94A - cont'd Geomorphic location of pond Toe Toe Toe Body Toe Head Body Body Head Toe Head Body Head Body Head Body Toe Body Toe Body Toe Body Body Head Toe Body Body Toe Body Head Body Body Pond Pond Slide Size Slide UTMs Slide Type # # (ha) 0.0241 696 10 10U 652580 6220750 Retrogressive rotational failure 0.0242 701 10 10U 652580 6220750 Retrogressive rotational failure 0.0243 764 224 10U 673168 6235959 Retrogressive rotational failure 0.0246 394 105 10U 607741 6250210 Compound failure 0.0246 464 122 10U 619929 6209531 Mobile flow 0.0251 325 90 10U 584584 6239849 Retrogressive rotational failure 0.0253 790 20 10U 679534 6219946 Rotational failure 0.0254 614 172 10U 641260 6254208 Rotational failure 0.0254 340 94 10U 586742 6231586 Rotational failure 0.0255 300 85 10U 612971 6237140 Retrogressive rotational failure 0.0255 32 25 10U 661743 6223747 Compound failure 0.0257 386 103 10U 589643 6282999 Rotational failure 0.0258 264 83 10U 611833 6238026 Retrogressive rotational failure 0.0259 758 219 10U 672065 6223288 Retrogressive rotational failure 0.0264 773 228 10U 676317 6244415 Retrogressive rotational failure 0.0265 529 144 10U 626024 6221319 Rotational failure 0.0266 260 3 10U 635265 6222327 Retrogressive rotational failure 0.0267 523 140 10U 634447 6231204 Retrogressive rotational failure 0.0268 40 29 10U 659264 6228605 Rotational failure 0.0271 656 189 10U 654871 6235759 Retrogressive rotational failure 0.0273 675 195 10U 655685 6231888 Rotational failure 0.0274 303 85 10U 612971 6237140 Retrogressive rotational failure 0.0276 706 10 10U 652580 6220750 Retrogressive rotational failure 0.0276 102 41 10U 647664 6238258 Retrogressive rotational failure 0.0277 274 83 10U 611833 6238026 Retrogressive rotational failure 0.0279 56 33 10U 657471 6230119 Compound failure 0.0281 689 200 10U 658361 6228323 Multi-level rotational failure 0.0283 544 3 10U 635265 6222327 Retrogressive rotational failure 0.0285 662 190 10U 653976 6235516 Retrogressive rotational failure 0.0285 780 232 10U 681946 6227934 Retrogressive rotational failure 0.0286 756 217 10U 662941 6224282 Retrogressive rotational failure 0.0287 413 110 10U 608193 6247991 Compound failure Pond location on slide Above small scarp (in debris) Above small scarp (in debris) Within/below headscarp (small, intact) Small scarp (in debris) Top of slide along west sidescarp Above small retrogressive scarp (intact) Base of headscarp (large, intact) Debris apron at base of new headscarp (large, intact) Debris apron Below small scarp (in debris) Below small scarp (intact) Debris apron Below small scarp (in debris) Below small scarp (intact) Within headscarp (small, intact) Just above small scarp (intact) Above small scarp, below headscarp (large, intact) Right above small scarp (intact) Debris apron Beside creek Above small scarp (intact) Below small scarp (in debris) Next to small scarp (in debris) Just below large scarp (intact) At base of large scarp (intact) Below large scarp (intact) Below small scarp (in debris) Below small scarp (in debris) Apron at top of slide below headscarp (large, intact) Base of slide next to creek Above small scarp (intact) Below small scarp (in debris) 285 Dormant Dormant Dormant Dormant Active Dormant Dormant Active Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Abandoned Dormant Abandoned Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Slide Activity Appendix 7 Landslide pond descriptions - Mapsheet 94A - cont'd Geomorphic location of pond Head Body Head Body Head Body Body Head Body Toe Body Body Body Body Head Toe Head Body Toe Toe Body Toe Body Toe Body Toe Toe Body Head Toe Body Body Pond Pond Slide Size Slide UTMs Slide Type # # (ha) 0.0288 589 47 10U 640425 6241038 Retrogressive rotational failure 0.0288 784 235 10U 685591 6226178 Multi-level rotational failure 0.0288 389 103 10U 589643 6282999 Rotational failure 0.0289 396 105 10U 607741 6250210 Compound failure 0.0289 125 57 10U 645882 6252696 Retrogressive rotational failure 0.0291 230 4 10U 637998 6220826 Multi-level rotational failure 0.0296 426 114 10U 607383 6243249 Retrogressive rotational failure 0.0296 757 218 10U 663556 6223926 Retrogressive rotational failure 0.0299 597 163 10U 640344 6243435 Multi-level rotational failure 0.0302 571 152 10U 637789 6222601 Retrogressive rotational failure 0.0304 524 142 10U 630879 6227122 Retrogressive rotational failure 0.0307 229 4 10U 637998 6220826 Multi-level rotational failure 0.0307 716 204 10U 654675 6219635 Retrogressive rotational failure 0.0307 58 33 10U 657471 6230119 Compound failure 0.0309 705 10 10U 652580 6220750 Retrogressive rotational failure 0.0309 772 227 10U 673362 6239998 Retrogressive rotational failure 0.031 499 136 10U 627698 6232647 Rotational failure 0.0316 646 186 10U 655856 6238250 Retrogressive rotational failure 0.0318 698 10 10U 652580 6220750 Retrogressive rotational failure 0.0319 359 97 10U 597665 6245826 Rotational failure 0.0321 625 178 10U 637541 6284374 Retrogressive rotational failure 0.0321 727 208 10U 655089 6218426 Retrogressive rotational failure 0.0321 322 90 10U 584584 6239849 Retrogressive rotational failure 0.0322 603 166 10U 641866 6247606 Retrogressive rotational failure 0.0327 491 133 10U 633983 6299243 Rotational failure 0.033 777 230 10U 673848 6240822 Rotational failure 0.0331 2 57 10U 645882 6252696 Retrogressive rotational failure 0.0334 406 107 10U 607530 6250024 Compound failure 0.0334 401 105 10U 607741 6250210 Compound failure 0.0335 465 122 10U 619929 6209531 Mobile flow 0.0338 124 57 10U 645882 6252696 Retrogressive rotational failure 0.0338 719 205 10U 653493 6218800 Multi-level rotational failure Pond location on slide Below small scarp (in debris) Base of large scarp (intact), in debris apron Debris apron near river Within small scarp (in debris) Above small scarp (in debris) Debris apron - near creek Below headscarp (large, intact) Below headscarp (large, intact), above small scarp (intact) Below headscarp (large, intact) Above small scarp (intact) Base of headscarp (small, intact) Debris apron Above small scarp (intact) Above small scarp (intact) Below small scarp (in debris) Within headscarp (small, intact) Above small scarp (intact), near river Base of small scarp (in debris), by creek Below small scarp (in debris) Within small scarp (in debris) Above small scarp (in debris) Above small scarp (intact) Just above small retrogressive scarp (intact) Below small scarp (intact) Debris apron Below headscarp (small, intact) Below small scarp (intact) Below headscarp (small, intact) Within small scarp (in debris) Top of slide along west sidescarp Below headscarp (large, intact) Base of headscarp (small, intact) 286 Dormant Dormant Dormant Dormant Active Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Active Dormant Dormant Active Active Dormant Slide Activity Appendix 7 Landslide pond descriptions - Mapsheet 94A - cont'd Geomorphic location of pond Body Toe Toe Body Body Toe Head Body Head Body Head Toe Body Body Body Head Body Toe Body Body Body Body Body Body Toe Body Toe Body Body Head Body Head Pond Pond Slide Size Slide UTMs Slide Type # # (ha) 0.034 190 10 10U 652580 6220750 Retrogressive rotational failure 0.0343 292 84 10U 612699 6237705 Retrogressive rotational failure 0.0346 731 14 10U 655144 6217752 Compound failure 0.0347 436 115 10U 606706 6242217 Rotational failure 0.035 275 83 10U 611833 6238026 Retrogressive rotational failure 0.0351 525 142 10U 630879 6227122 Retrogressive rotational failure 0.0354 741 212 10U 656103 6216700 Retrogressive rotational failure 0.0356 227 4 10U 637998 6220826 Multi-level rotational failure 0.0356 455 122 10U 619929 6209531 Mobile flow 0.0357 754 215 10U 664950 6210554 Multi-level rotational failure 0.0357 17 57 10U 645882 6252696 Retrogressive rotational failure 0.0361 738 15 10U 655647 6215120 Retrogressive rotational failure 0.0363 692 201 10U 655452 6222977 Retrogressive rotational failure 0.0365 279 83 10U 611833 6238026 Retrogressive rotational failure 0.0367 513 141 10U 635411 6231647 Retrogressive rotational failure 0.0375 510 140 10U 634447 6231204 Retrogressive rotational failure 0.0375 676 195 10U 655685 6231888 Rotational failure 0.0379 621 176 10U 638057 6276259 Retrogressive rotational failure 0.0382 707 10 10U 652580 6220750 Retrogressive rotational failure 0.0384 488 132 10U 633416 6298822 Rotational failure 0.0384 541 3 10U 635265 6222327 Retrogressive rotational failure 0.0385 313 89 10U 583121 6249823 Multi-level rotational failure 0.0385 46 71 10U 660889 6229983 Shallow retrogressive failure 0.0387 181 17 10U 672017 6212944 Rotational failure 0.0389 549 145 10U 634428 6225821 Multi-level rotational failure 0.0389 717 204 10U 654675 6219635 Retrogressive rotational failure 0.0397 650 188 10U 655498 6235114 Retrogressive rotational failure 0.0398 585 47 10U 640425 6241038 Retrogressive rotational failure 0.04 583 161 10U 639518 6240605 Retrogressive rotational failure 0.0402 605 53 10U 641413 6249461 Retrogressive rotational failure 0.0405 3 57 10U 645882 6252696 Retrogressive rotational failure 0.0405 441 117 10U 605801 6243804 Rotational failure Pond location on slide Below small scarp (intact) Just below small scarp (intact) Below small scarp (in debris) Near creek/river At base of large scarp (intact) Base of headscarp (small, intact) Bottom of slide, by creek Debris apron Near top of slide Base of headscarp (small, intact) Just above small scarp (in debris) Above/within small scarp (in debris) Bottom of slide Toe Flat area midway down slide Below small scarp (in debris) Below headscarp (small, intact) Below headscarp (large, intact) Below small scarp (in debris) Below large scarp (intact), near river Below small scarp (in debris) Debris apron next to river Below headscarp (small, intact) Below small scarp (intact) Debris apron Above small scarp (intact), below headscarp (small, intact) Below headscarp (large, intact) Halfway down slide Below small scarp (intact), near river Below headscarp (large, intact) Below small scarp (intact) Below headscarp (large, intact), above small scarp (in debris) 287 Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Active Dormant Active Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Active Dormant Slide Activity Appendix 7 Landslide pond descriptions - Mapsheet 94A - cont'd Geomorphic location of pond Toe Body Body Toe Body Head Toe Toe Body Head Body Body Toe Toe Toe Body Body Head Body Toe Body Toe Body Body Toe Head Head Body Toe Body Toe Body Pond Pond Slide Size Slide UTMs Slide Type # # (ha) 0.0406 307 87 10U 580867 6250762 Compound failure 0.0406 201 13 10U 656024 6217434 Compound failure 0.0409 736 210 10U 655750 6214657 Rotational failure 0.0414 60 36 10U 658291 6231793 Retrogressive rotational failure 0.0414 22 57 10U 645882 6252696 Retrogressive rotational failure 0.0415 129 64 10U 641133 6264748 Retrogressive rotational failure 0.0417 688 200 10U 658361 6228323 Multi-level rotational failure 0.0418 598 164 10U 640997 6242676 Rotational failure 0.0419 284 83 10U 611833 6238026 Retrogressive rotational failure 0.0419 570 152 10U 637789 6222601 Retrogressive rotational failure 0.0421 336 92 10U 584580 6235119 Rotational failure 0.0422 788 236 10U 675638 6221766 Multi-level rotational failure 0.0425 101 45 10U 645103 6237317 Retrogressive rotational failure 0.0427 504 137 10U 628079 6232225 Multi-level rotational failure 0.0428 199 12 10U 656272 6218290 Rotational failure 0.0429 306 87 10U 580867 6250762 Compound failure 0.0435 81 40 10U 648407 6238441 Multi-level rotational failure 0.0435 568 154 10U 636856 6222710 Compound failure 0.0436 387 103 10U 589643 6282999 Rotational failure 0.0437 141 74 10U 636436 6241818 Compound failure 0.0438 626 178 10U 637541 6284374 Retrogressive rotational failure 0.0439 39 28 10U 660309 6228226 Compound failure 0.044 543 3 10U 635265 6222327 Retrogressive rotational failure 0.0442 35 27 10U 661452 6225535 Compound failure 0.0443 85 41 10U 647664 6238258 Retrogressive rotational failure 0.0446 785 235 10U 685591 6226178 Multi-level rotational failure 0.0447 791 238 10U 679053 6219617 Rotational failure 0.0451 492 133 10U 633983 6299243 Rotational failure 0.0452 397 105 10U 607741 6250210 Compound failure 0.0452 712 10 10U 652580 6220750 Retrogressive rotational failure 0.0456 461 122 10U 619929 6209531 Mobile flow 0.0456 182 17 10U 672017 6212944 Rotational failure Pond location on slide Just below headscarp (small, intact) Below small scarp (in debris) Below headscarp (small, intact), next to north sidescarp Toe Below small scarp (in debris) Below headscarp (large, intact) Above small scarp (intact) Above small scarp (in debris) At bottom of slide Base of small scarp (intact) Within small scarp (in debris) Debris apron at base of slide Just above small scarp (intact) Above small scarp (intact), near east sidescarp Above small scarp (in debris) Just below headscarp (small, intact) Debris apron below large scarp (intact) Above small scarp (intact), near east sidescarp Debris apron right by river Below headscarp (large, intact) Below small scarp (intact), near east sidescarp Toe Below small scarp (in debris) Body Below small scarp (intact) Base of large scarp (intact), in debris apron Debris apron Debris apron Debris apron near river Next to small scarp (in debris) Top of slide along west sidescarp Below small scarp (intact) 288 Dormant Dormant Dormant Dormant Active Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Active Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Active Dormant Slide Activity Appendix 7 Landslide pond descriptions - Mapsheet 94A - cont'd Geomorphic location of pond Head Body Head Toe Toe Head Body Body Toe Body Body Toe Body Body Body Head Toe Body Toe Body Toe Toe Body Body Body Toe Body Toe Toe Head Head Body Pond Pond Slide Size Slide UTMs Slide Type # # (ha) 0.0457 721 207 10U 655366 6218822 Retrogressive rotational failure 0.0458 159 63 10U 656420 6272207 Rotational failure 0.0462 740 211 10U 655836 6216390 Retrogressive rotational failure 0.0462 708 10 10U 652580 6220750 Retrogressive rotational failure 0.0463 192 10 10U 652580 6220750 Retrogressive rotational failure 0.0465 453 122 10U 619929 6209531 Mobile flow 0.0466 312 88 10U 582374 6251325 Rotational failure 0.047 281 83 10U 611833 6238026 Retrogressive rotational failure 0.0472 223 5 10U 637848 6220421 Retrogressive rotational failure 0.0473 162 22 10U 668246 6222087 Compound failure 0.0473 768 227 10U 673362 6239998 Retrogressive rotational failure 0.0473 390 103 10U 589643 6282999 Rotational failure 0.0474 700 10 10U 652580 6220750 Retrogressive rotational failure 0.0474 152 61 10U 648617 6262829 Retrogressive rotational failure 0.0474 553 146 10U 635074 6225748 Multi-level rotational failure 0.0476 792 89 10U 583121 6249823 Multi-level rotational failure 0.0478 252 1 10U 631931 6221078 Retrogressive rotational failure 0.048 622 176 10U 638057 6276259 Retrogressive rotational failure 0.0485 188 10 10U 652580 6220750 Retrogressive rotational failure 0.0488 442 118 10U 605649 6243226 Rotational failure 0.0491 388 103 10U 589643 6282999 Rotational failure 0.0494 268 83 10U 611833 6238026 Retrogressive rotational failure 0.0498 486 130 10U 633609 6300494 Compound failure 0.0498 225 5 10U 637848 6220421 Retrogressive rotational failure 0.0498 534 1 10U 631931 6221078 Retrogressive rotational failure 0.0499 711 10 10U 652580 6220750 Retrogressive rotational failure 0.0499 454 122 10U 619929 6209531 Mobile flow 0.05 26 60 10U 647207 6252566 Rotational failure 0.0501 120 54 10U 642211 6250812 Rotational failure 0.0506 651 188 10U 655498 6235114 Retrogressive rotational failure 0.0507 195 11 10U 656217 6219302 Retrogressive rotational failure 0.0508 361 98 10U 598431 6246464 Compound failure Pond location on slide Base of headscarp (small, intact) Below small scarp (in debris) Bottom of slide, near west sidescarp Below small scarp (in debris) Below small scarp (intact) Near base of slide Within debris of small scarp (in debris) Toe Just above small scarp (in debris) Above small scarp (intact) Base of headscarp (small, intact) Debris apron near river Below small scarp (in debris) In fresh material below small scarp (in debris) Above small scarp (in debris) Debris apron next to river Above backtilt Within small scarp (intact) Below small scarp (intact) Just above small scarp (intact) Debris apron Above large scarp (intact), below small scarp (in debris) Within small scarp (intact) Below headscarp (small, intact) Below small scarp (in debris) Below headscarp (small, intact) Near centre of slide Debris apron In debris apron at base of headscarp (large, intact) Near creek Just below small scarp (in debris) Just below headscarp (small, intact) 289 Dormant Dormant Dormant Dormant Dormant Active Dormant Dormant Active Dormant Dormant Dormant Dormant Active Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Active Dormant Dormant Active Dormant Dormant Dormant Active Dormant Slide Activity Appendix 7 Landslide pond descriptions - Mapsheet 94A - cont'd Geomorphic location of pond Head Toe Toe Body Toe Body Toe Toe Body Body Head Toe Body Toe Head Toe Body Body Toe Body Body Body Body Head Body Head Body Toe Body Toe Body Head Pond Pond Slide Size Slide UTMs Slide Type # # (ha) 0.0508 673 194 10U 656093 6232341 Retrogressive rotational failure 0.0512 431 114 10U 607383 6243249 Retrogressive rotational failure 0.0513 265 83 10U 611833 6238026 Retrogressive rotational failure 0.0516 769 227 10U 673362 6239998 Retrogressive rotational failure 0.0522 205 14 10U 655144 6217752 Compound failure 0.0525 726 208 10U 655089 6218426 Retrogressive rotational failure 0.0528 149 76 10U 633480 6244644 Retrogressive rotational failure 0.0528 775 230 10U 673848 6240822 Rotational failure 0.0529 729 14 10U 655144 6217752 Compound failure 0.053 564 4 10U 637998 6220826 Multi-level rotational failure 0.0531 135 70 10U 631913 6311160 Retrogressive rotational failure 0.0533 269 83 10U 611833 6238026 Retrogressive rotational failure 0.0535 89 43 10U 646755 6238196 Retrogressive rotational failure 0.0539 463 122 10U 619929 6209531 Mobile flow 0.0539 507 139 10U 633271 6230890 Compound failure 0.054 528 143 10U 624881 6220596 Retrogressive rotational failure 0.0547 118 53 10U 641413 6249461 Retrogressive rotational failure 0.0547 285 83 10U 611833 6238026 Retrogressive rotational failure 0.0548 83 40 10U 648407 6238441 Multi-level rotational failure 0.0551 540 3 10U 635265 6222327 Retrogressive rotational failure 0.0551 443 119 10U 599356 6238157 Multi-level rotational failure 0.0552 439 116 10U 606417 6242052 Rotational failure 0.056 508 140 10U 634447 6231204 Retrogressive rotational failure 0.0563 311 88 10U 582374 6251325 Rotational failure 0.0565 642 186 10U 655856 6238250 Retrogressive rotational failure 0.0565 734 210 10U 655750 6214657 Rotational failure 0.0567 697 10 10U 652580 6220750 Retrogressive rotational failure 0.0569 458 122 10U 619929 6209531 Mobile flow 0.0571 314 89 10U 583121 6249823 Multi-level rotational failure 0.0576 364 99 10U 598103 6246815 Rotational failure 0.0578 293 84 10U 612699 6237705 Retrogressive rotational failure 0.0579 95 45 10U 645103 6237317 Retrogressive rotational failure Pond location on slide Below headscarp (large, intact) Below large scarp (intact) Above large scarp (intact) Base of headscarp (small, intact) Above small scarp (in debris) Base of headscarp (small, intact) Just above small scarp (in debris) Within headscarp (small, intact) Above small scarp (in debris) Base of small scarp (intact) Below headscarp (small, intact) At base of headscarp (large, intact) Just below small scarp (intact) Top of slide near west sidescarp Above small scarp (in debris) Below small scarp (intact), near north sidescarp Just below headscarp (large, intact) At bottom of slide Just below large scarp (intact) Above small scarp (in debris) Base of large scarp (intact) Debris apron Above small scarp (intact) Just below small scarp (in debris) Below headscarp (large, intact) Just above small scarp (in debris) Above small scarp (in debris) Near top of slide Debris apron next to river At base of headscarp (large, intact), by sidescarp Within small scarp (in debris) Below small scarp (intact) 290 Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Active Active Dormant Dormant Dormant Dormant Active Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Active Dormant Dormant Dormant Dormant Slide Activity Appendix 7 Landslide pond descriptions - Mapsheet 94A - cont'd Geomorphic location of pond Body Body Body Head Body Head Body Head Body Body Body Body Body Head Body Toe Body Toe Body Body Body Toe Body Toe Head Body Head Body Toe Head Body Toe Pond Pond Slide Size Slide UTMs Slide Type # # (ha) 0.0587 674 194 10U 656093 6232341 Retrogressive rotational failure 0.0588 451 122 10U 619929 6209531 Mobile flow 0.0589 645 186 10U 655856 6238250 Retrogressive rotational failure 0.0593 337 92 10U 584580 6235119 Rotational failure 0.0594 53 71 10U 660889 6229983 Shallow retrogressive failure 0.0598 733 209 10U 655227 6215278 Retrogressive rotational failure 0.06 220 4 10U 637998 6220826 Multi-level rotational failure 0.0603 632 181 10U 654030 6269428 Rotational failure 0.0608 298 85 10U 612971 6237140 Retrogressive rotational failure 0.061 41 31 10U 657777 6229034 Compound failure 0.0611 143 75 10U 633471 6243680 Retrogressive rotational failure 0.0621 466 85 10U 612971 6237140 Retrogressive rotational failure 0.0622 79 40 10U 648407 6238441 Multi-level rotational failure 0.0628 702 10 10U 652580 6220750 Retrogressive rotational failure 0.0629 782 234 10U 684625 6226289 Retrogressive rotational failure 0.0633 448 122 10U 619929 6209531 Mobile flow 0.0644 55 72 10U 659938 6229825 Retrogressive rotational failure 0.065 339 94 10U 586742 6231586 Rotational failure 0.0651 728 14 10U 655144 6217752 Compound failure 0.0651 759 220 10U 673724 6221673 Multi-level rotational failure 0.0656 434 114 10U 607383 6243249 Retrogressive rotational failure 0.0658 383 101 10U 598608 6247865 Compound failure 0.066 624 177 10U 638157 6284497 Rotational failure 0.0662 575 157 10U 643272 6221186 Retrogressive rotational failure 0.0665 310 87 10U 580867 6250762 Compound failure 0.0667 119 53 10U 641413 6249461 Retrogressive rotational failure 0.0669 765 222 10U 672726 6234762 Retrogressive rotational failure 0.0673 34 27 10U 661452 6225535 Compound failure 0.0679 45 71 10U 660889 6229983 Shallow retrogressive failure 0.068 217 4 10U 637998 6220826 Multi-level rotational failure 0.0686 384 102 10U 596707 6242884 Rotational failure 0.069 222 5 10U 637848 6220421 Retrogressive rotational failure Pond location on slide Just below small scarp (in debris) Centre of slide Below large headscarp (intact) Just below headscarp (large, intact) Just below small scarp (intact) Base of small scarp (in debris) Below small scarp (intact) Bottom of slide Below small scarp (in debris) Below small scarp (intact) Just below small scarp (intact) Below small scarp (in debris) Debris apron Above small scarp (in debris) Below/within small scarp (intact) Near base of slide by west sidescarp Near base of headscarp (large), above small scarp (in debris) At base of headscarp (large, intact) Bottom of slide, near north sidescarp Above small scarp (intact) Next to south sidescarp Debris apron below headscarp (small, intact) Base of small scarp (intact) Below large scarp (intact) Debris apron, just above large scarp (small, intact) Below headscarp (large, intact) Next to creek Body Below headscarp (small, intact) Below headscarp (large, intact) Above small scarp (in debris) Below headscarp (small, intact) 291 Dormant Active Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Active Dormant Active Dormant Dormant Active Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Active Slide Activity Appendix 7 Landslide pond descriptions - Mapsheet 94A - cont'd Geomorphic location of pond Body Body Head Body Toe Body Body Toe Toe Toe Body Body Toe Body Body Body Body Head Toe Body Body Body Toe Head Body Body Toe Body Body Head Body Head Pond Pond Slide Size Slide UTMs Slide Type # # (ha) 0.0691 24 59 10U 646337 6252998 Retrogressive rotational failure 0.0702 196 11 10U 656217 6219302 Retrogressive rotational failure 0.0705 743 212 10U 656103 6216700 Retrogressive rotational failure 0.0707 301 85 10U 612971 6237140 Retrogressive rotational failure 0.0709 559 149 10U 635558 6216354 Rotational failure 0.0711 180 17 10U 672017 6212944 Rotational failure 0.0721 324 90 10U 584584 6239849 Retrogressive rotational failure 0.0723 569 152 10U 637789 6222601 Retrogressive rotational failure 0.0726 435 115 10U 606706 6242217 Rotational failure 0.0732 27 60 10U 647207 6252566 Rotational failure 0.0733 714 202 10U 654150 6220760 Retrogressive rotational failure 0.0739 108 48 10U 641990 6244350 Rotational failure 0.0747 33 26 10U 661704 6225287 Shallow retrogressive failure 0.075 693 201 10U 655452 6222977 Retrogressive rotational failure 0.0751 793 90 10U 584584 6239849 Retrogressive rotational failure 0.0754 405 106 10U 607981 6249961 Rotational failure 0.0755 685 33 10U 657471 6230119 Compound failure 0.0767 670 194 10U 656093 6232341 Retrogressive rotational failure 0.0771 93 45 10U 645103 6237317 Retrogressive rotational failure 0.0777 643 186 10U 655856 6238250 Retrogressive rotational failure 0.0779 710 10 10U 652580 6220750 Retrogressive rotational failure 0.0782 433 114 10U 607383 6243249 Retrogressive rotational failure 0.079 140 74 10U 636436 6241818 Compound failure 0.0793 91 43 10U 646755 6238196 Retrogressive rotational failure 0.0798 636 182 10U 653926 6268718 Retrogressive rotational failure 0.0809 54 71 10U 660889 6229983 Shallow retrogressive failure 0.0816 13 58 10U 646995 6252365 Compound failure 0.0816 438 115 10U 606706 6242217 Rotational failure 0.0817 663 39 10U 651511 6236995 Rotational failure 0.0821 437 115 10U 606706 6242217 Rotational failure 0.0823 699 10 10U 652580 6220750 Retrogressive rotational failure 0.0836 355 96 10U 597364 6244753 Compound failure Pond location on slide Toe Just below small scarp (in debris) Above small scarp (intact) Below small scarp (in debris) Within small scarp (in debris) Within headscarp (small, intact) Just above small retrogressive scarp (intact) Base of headscarp (small, intact) Above small scarp (intact) Debris apron Within small scarp (intact) At base of headscarp (large, intact) In debris apron Within small scarp (in debris) Just above small retrogressive scarp (intact) Debris apron near river Above large scarp (intact), near river Below headscarp (large, intact) Below small scarp (intact) Base of headscarp (large, intact) Below headscarp (small, intact) Above small scarp (intact) At base of headscarp (large, intact) Just below small scarp (intact) Just above small scarp (intact), near river Just above small scarp (intact) Below small scarp (in debris) Near west sidescarp Debris apron Below headscarp (large, intact) Below small scarp (in debris) Just below small scarp (in debris) 292 Dormant Active Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Active Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Slide Activity Appendix 7 Landslide pond descriptions - Mapsheet 94A - cont'd Geomorphic location of pond Toe Body Body Toe Body Head Body Head Body Toe Toe Head Toe Toe Body Body Body Head Toe Head Head Body Body Toe Toe Toe Toe Body Toe Body Body Body Pond Pond Slide Size Slide UTMs Slide Type # # (ha) 0.0837 288 83 10U 611833 6238026 Retrogressive rotational failure 0.086 444 120 10U 603024 6239487 Rotational failure 0.0863 601 50 10U 641564 6246268 Compound failure 0.0868 151 78 10U 637889 6241550 Rotational failure 0.0874 75 44 10U 646758 6237400 Retrogressive rotational failure 0.0874 760 221 10U 670755 6236278 Rotational failure 0.0876 783 234 10U 684625 6226289 Retrogressive rotational failure 0.0877 385 103 10U 589643 6282999 Rotational failure 0.0881 407 107 10U 607530 6250024 Compound failure 0.089 343 94 10U 586742 6231586 Rotational failure 0.0915 123 55 10U 641685 6251522 Multi-level rotational failure 0.0923 779 231 10U 680412 6226004 Multi-level rotational failure 0.0927 391 103 10U 589643 6282999 Rotational failure 0.0928 287 83 10U 611833 6238026 Retrogressive rotational failure 0.093 161 22 10U 668246 6222087 Compound failure 0.0931 11 58 10U 646995 6252365 Compound failure 0.0942 753 215 10U 664950 6210554 Multi-level rotational failure 0.0944 709 10 10U 652580 6220750 Retrogressive rotational failure 0.0947 539 3 10U 635265 6222327 Retrogressive rotational failure 0.095 321 90 10U 584584 6239849 Retrogressive rotational failure 0.0958 273 83 10U 611833 6238026 Retrogressive rotational failure 0.0976 187 10 10U 652580 6220750 Retrogressive rotational failure 0.0995 200 13 10U 656024 6217434 Compound failure 0.0996 29 23 10U 662280 6223095 Rotational failure 0.1017 144 75 10U 633471 6243680 Retrogressive rotational failure 0.1019 563 4 10U 637998 6220826 Multi-level rotational failure 0.1028 745 212 10U 656103 6216700 Retrogressive rotational railure 0.1031 690 30 10U 658107 6228756 Compound failure 0.1033 197 12 10U 656272 6218290 Rotational failure 0.1035 672 194 10U 656093 6232341 Retrogressive rotational failure 0.1045 122 55 10U 641685 6251522 Multi-level rotational failure 0.1045 595 162 10U 639763 6241926 Retrogressive rotational failure Pond location on slide At bottom of slide Just below headscarp (large, intact) Within small scarp (in debris) Just below headscarp (small) Below small scarp (in debris) Base of headscarp (large, intact) Below small scarp (intact) At base of headscarp (small, intact) Below headscarp (small, intact) near south sidescarp Debris apron Debris apron Debris apron at base of large scarp (intact) Debris apron near river At bottom of slide Below small scarp (intact) Below headscarp (small, intact) Base of headscarp (small, intact) Below small scarp (in debris) Just below small scarp (intact) Just above small retrogressive scarp (intact) Below large scarp (intact) Below small scarp (intact) Above small scarp (in debris), just below headscarp (small, intact) Above debris apron Below small scarp (intact), next to small scarp (in debris) Below small scarp (intact) Below headscarp (large, intact) Below small scarp (in debris) Above small scarp (in debris) Just below small scarp (in debris) Debris apron Below small scarp (in debris) 293 Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Abandoned Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Active Active Dormant Dormant Dormant Dormant Dormant Abandoned Dormant Slide Activity Appendix 7 Landslide pond descriptions - Mapsheet 94A - cont'd Geomorphic location of pond Toe Head Body Head Body Head Body Head Body Body Toe Toe Toe Toe Body Head Head Body Body Body Body Toe Head Body Body Body Body Toe Body Body Toe Body Pond Pond Slide Size Slide UTMs Slide Type # # (ha) 0.1049 794 97 10U 597665 6245826 Rotational failure 0.1055 648 187 10U 655524 6236858 Multi-level rotational failure 0.1057 215 86 10U 639774 6220198 Retrogressive rotational failure 0.1074 581 161 10U 639518 6240605 Retrogressive rotational failure 0.1086 612 172 10U 641260 6254208 Rotational failure 0.1091 462 122 10U 619929 6209531 Mobile flow 0.1094 786 235 10U 685591 6226178 Multi-level rotational failure 0.1096 482 128 10U 632934 6302871 Rotational failure 0.1101 216 4 10U 637998 6220826 Multi-level rotational failure 0.1103 226 4 10U 637998 6220826 Multi-level rotational failure 0.1106 654 189 10U 654871 6235759 Retrogressive rotational failure 0.1112 752 215 10U 664950 6210554 Multi-level rotational failure 0.1116 778 227 10U 673362 6239998 Retrogressive rotational failure 0.1121 750 214 10U 664400 6210645 Multi-level rotational failure 0.1121 776 230 10U 673848 6240822 Rotational failure 0.1129 565 152 10U 637789 6222601 Retrogressive rotational failure 0.1136 6 59 10U 646337 6252998 Retrogressive rotational failure 0.1139 577 158 10U 643429 6220210 Rotational failure 0.1152 720 206 10U 655413 6219392 Retrogressive rotational failure 0.1161 323 90 10U 584584 6239849 Retrogressive rotational failure 0.117 762 81 10U 673096 6235267 Retrogressive rotational failure 0.1173 590 47 10U 640425 6241038 Retrogressive rotational failure 0.1194 15 57 10U 645882 6252696 Retrogressive rotational failure 0.1195 449 122 10U 619929 6209531 Mobile flow 0.1205 92 45 10U 645103 6237317 Retrogressive rotational failure 0.1218 145 75 10U 633471 6243680 Retrogressive rotational failure 0.1223 19 57 10U 645882 6252696 Retrogressive rotational failure 0.1229 256 2 10U 633364 6222821 Compound failure 0.1233 142 74 10U 636436 6241818 Compound failure 0.1233 432 114 10U 607383 6243249 Retrogressive rotational failure 0.1235 331 91 10U 585686 6240356 Compound failure 0.1238 12 57 10U 645882 6252696 Retrogressive rotational failure Pond location on slide At base of headscarp (large, intact) Below headscarp (large, intact) Below small scarp (intact) Below small scarp (in debris), near river Debris apron beside creek Top of slide near west sidescarp Base of large scarp (intact), in debris apron Base of headscarp (large, intact) Below headscarp (large, intact), above small scarp (intact) Debris apron Below headscarp (large, intact) Base of headscarp (small, intact) Base of slide, near creek Below headscarp (small, intact) Within headscarp (small, intact) Above small scarp (intact), next to east sidescarp Below small scarp (in debris) Below small scarp (in debris), by north sidescarp Below headscarp (small, intact) Below small scarp (in debris) Above small scarp (intact), near creek Halfway down slide Just above small scarp (in debris) Near base of slide by west sidescarp Within large scarp (intact), below headscarp (small, intact) Just below small scarp (intact) Below small scarp (in debris) Below small scarp (in debris) Below small scarp (intact) Below large scarp (intact) Just below small scarp (intact) Just above small scarp (in debris) 294 Dormant Dormant Dormant Dormant Active Active Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Active Active Dormant Active Active Abandoned Dormant Dormant Dormant Active Slide Activity Appendix 7 Landslide pond descriptions - Mapsheet 94A - cont'd Geomorphic location of pond Head Head Head Toe Toe Head Toe Body Head Toe Body Head Toe Head Head Body Body Toe Head Toe Body Body Body Body Head Body Body Toe Body Body Body Body Pond Pond Slide Size Slide UTMs Slide Type # # (ha) 0.1244 467 123 10U 615696 6235258 Multi-level rotational failure 0.1269 573 156 10U 643924 6221112 Rotational failure 0.1275 479 127 10U 632593 6304450 Rotational failure 0.1281 354 96 10U 597364 6244753 Compound failure 0.1284 412 109 10U 608185 6248414 Compound failure 0.131 107 48 10U 641990 6244350 Rotational failure 0.131 127 56 10U 644848 6253033 Retrogressive rotational failure 0.1311 671 194 10U 656093 6232341 Retrogressive rotational failure 0.1317 658 189 10U 654871 6235759 Retrogressive rotational failure 0.1324 212 9 10U 639115 6218246 Rotational failure 0.1338 501 137 10U 628079 6232225 Multi-level rotational failure 0.137 139 73 10U 634577 6242061 Compound failure 0.1383 537 3 10U 635265 6222327 Retrogressive rotational failure 0.1396 358 97 10U 597665 6245826 Rotational failure 0.1409 666 38 10U 652244 6236095 Retrogressive rotational failure 0.1415 771 227 10U 673362 6239998 Retrogressive rotational failure 0.1417 761 222 10U 672726 6234762 Retrogressive rotational failure 0.1418 255 1 10U 631931 6221078 Retrogressive rotational failure 0.1419 228 4 10U 637998 6220826 Multi-level rotational failure 0.1423 665 192 10U 649524 6238522 Rotational failure 0.1425 277 83 10U 611833 6238026 Retrogressive rotational failure 0.1431 224 5 10U 637848 6220421 Retrogressive rotational failure 0.1437 604 167 10U 642365 6247706 Rotational failure 0.1439 578 159 10U 649510 6220866 Rotational failure 0.1441 14 57 10U 645882 6252696 Retrogressive rotational failure 0.1441 580 160 10U 648168 6224229 Compound failure 0.1442 116 53 10U 641413 6249461 Retrogressive rotational failure 0.1448 179 19 10U 675077 6219231 Rotational failure 0.1452 450 122 10U 619929 6209531 Mobile flow 0.1457 591 47 10U 640425 6241038 Retrogressive rotational failure 0.1467 7 59 10U 646337 6252998 Retrogressive rotational failure 0.1476 715 203 10U 654136 6220177 Retrogressive rotational failure 0.1481 703 10 10U 652580 6220750 Retrogressive rotational failure Debris apron near river Within small scarp (in debris) Within small scarp (intact) Within small scarp (in debris) Debris apron near river/creek At base of headscarp (large, intact) Below small scarp Below small scarp (in debris) Above small scarp (intact) Below small scarp (intact) Below small scarp (intact), by southwest sidescarp Below small scarp (in debris) Above small scarp (intact) At base of headscarp (large, intact) Next to river at bottom of slide Base of headscarp (small, intact) Above small scarp (in debris), near creek Above small scarp (intact) Debris apron Below headscarp (small, intact) Below small scarp (in debris) Between 2 small scarps (in debris) Bottom of slide, next to creek Next to west side scarp, near base of slide At base of headscarp (large, intact) Above small scarp (in debris) At base of headscarp (large, intact) Debris apron Centre of slide Below, to the right of small scarp (in debris) Below headscarp (large, intact) Above small scarp (in debris), near south sidescarp Below small scarp (in debris) Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Active Dormant Dormant Active Dormant Dormant Dormant Active Dormant Dormant Dormant Dormant 295 Pond location on slide Slide Activity Appendix 7 Landslide pond descriptions - Mapsheet 94A - cont'd Geomorphic location of pond Toe Body Body Body Toe Head Toe Body Body Toe Body Body Body Head Toe Head Toe Body Toe Body Body Body Toe Toe Head Body Body Body Body Body Body Toe Body Pond Pond Slide Size Slide UTMs Slide Type # # (ha) 0.1507 620 176 10U 638057 6276259 Retrogressive rotational failure 0.1518 94 45 10U 645103 6237317 Retrogressive rotational failure 0.1518 97 45 10U 645103 6237317 Retrogressive rotational failure 0.1534 189 10 10U 652580 6220750 Retrogressive rotational failure 0.1542 547 145 10U 634428 6225821 Multi-level rotational failure 0.1546 221 4 10U 637998 6220826 Multi-level rotational failure 0.1546 198 12 10U 656272 6218290 Rotational failure 0.1557 290 83 10U 611833 6238026 Retrogressive rotational failure 0.1561 138 73 10U 634577 6242061 Compound failure 0.1578 356 96 10U 597364 6244753 Compound failure 0.1591 291 83 10U 611833 6238026 Retrogressive rotational failure 0.1597 247 1 10U 631931 6221078 Retrogressive rotational failure 0.1607 551 145 10U 634428 6225821 Multi-level rotational failure 0.1614 735 15 10U 655647 6215120 Retrogressive rotational failure 0.1616 579 159 10U 649510 6220866 Rotational failure 0.1622 744 212 10U 656103 6216700 Retrogressive rotational failure 0.1634 684 33 10U 657471 6230119 Compound failure 0.1639 572 155 10U 644195 6222091 Multi-level rotational failure 0.1664 496 75 10U 633471 6243680 Retrogressive rotational failure 0.1668 746 213 10U 659241 6219068 Retrogressive rotational failure 0.1673 567 154 10U 636856 6222710 Compound failure 0.1674 755 216 10U 663258 6218654 Multi-level rotational failure 0.1689 1 15 10U 655647 6215120 Retrogressive rotational failure 0.1718 42 30 10U 658107 6228756 Compound failure 0.1726 242 1 10U 631931 6221078 Retrogressive rotational failure 0.1729 259 3 10U 635265 6222327 Retrogressive rotational failure 0.1733 249 1 10U 631931 6221078 Retrogressive rotational failure 0.1735 669 193 10U 657047 6232296 Retrogressive rotational failure 0.1761 206 16 10U 655340 6214299 Rotational failure 0.1761 276 83 10U 611833 6238026 Retrogressive rotational failure 0.1765 546 145 10U 634428 6225821 Multi-level rotational failure 0.1811 751 214 10U 664400 6210645 Multi-level rotational failure Pond location on slide Base of headscarp (large, intact) Below small scarp (intact) Below small scarp (intact) Below small scarp (intact) Debris apron Below small scarp (intact) Just below headscarp (small, intact) At bottom of slide Next to small scarp (intact) Next to north sidescarp At bottom of slide Backtilt below small scarp (intact) Debris apron Below headscarp (small, intact), along south sidescarp Above small scarp (intact) Below headscarp (large, intact) Below large scarp (intact), near river Below small scarp (intact) Below small scarp (intact) Below small scarp (intact) Below small scarp (intact), near east sidescarp Above small scarp (intact), near river Just below headscarp (small, intact) Next to small scarp (in debris) Below small scarp (intact) Above small scarp, below headscarp (large, intact) Next to small scarp (in debris) Below headscarp (large, intact) On headscarp (small, intact) Below small scarp (in debris) Debris apron Within headscarp (small, intact) 296 Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Abandoned Dormant Abandoned Dormant Dormant Dormant Dormant Dormant Dormant Slide Activity Appendix 7 Landslide pond descriptions - Mapsheet 94A - cont'd Geomorphic location of pond Head Toe Toe Toe Toe Body Head Toe Toe Body Toe Body Toe Head Body Head Toe Body Body Toe Body Toe Head Toe Body Head Body Head Head Body Toe Head Pond Pond Slide Size Slide UTMs Slide Type # # (ha) 0.1825 657 189 10U 654871 6235759 Retrogressive rotational failure 0.1828 121 55 10U 641685 6251522 Multi-level rotational failure 0.1841 76 44 10U 646758 6237400 Retrogressive rotational failure 0.1863 117 53 10U 641413 6249461 Retrogressive rotational failure 0.1874 596 163 10U 640344 6243435 Multi-level rotational failure 0.189 243 1 10U 631931 6221078 Retrogressive rotational failure 0.1897 704 10 10U 652580 6220750 Retrogressive rotational failure 0.1905 261 3 10U 635265 6222327 Retrogressive rotational failure 0.191 175 20 10U 679534 6219946 Rotational failure 0.1911 713 10 10U 652580 6220750 Retrogressive rotational failure 0.1937 113 50 10U 641564 6246268 Compound failure 0.197 158 63 10U 656420 6272207 Rotational failure 0.1975 115 52 10U 641810 6248528 Rotational failure 0.1978 251 1 10U 631931 6221078 Retrogressive rotational failure 0.2006 747 213 10U 659241 6219068 Retrogressive rotational failure 0.2009 640 185 10U 656213 6239712 Rotational failure 0.2012 137 73 10U 634577 6242061 Compound failure 0.2022 241 1 10U 631931 6221078 Retrogressive rotational failure 0.2033 789 237 10U 680337 6220929 Retrogressive rotational failure 0.209 299 85 10U 612971 6237140 Retrogressive rotational failure 0.2105 599 48 10U 641990 6244350 Rotational failure 0.2115 258 3 10U 635265 6222327 Retrogressive rotational failure 0.2182 594 162 10U 639763 6241926 Retrogressive otational failure 0.222 574 157 10U 643272 6221186 Retrogressive rotational failure 0.2223 428 114 10U 607383 6243249 Retrogressive rotational failure 0.2225 516 141 10U 635411 6231647 Retrogressive rotational failure 0.2228 533 1 10U 631931 6221078 Retrogressive rotational failure 0.2237 576 157 10U 643272 6221186 Retrogressive rotational failure 0.2251 618 174 10U 642740 6258340 Retrogressive rotational failure 0.2332 240 1 10U 631931 6221078 Retrogressive rotational failure 0.234 136 70 10U 631913 6311160 Retrogressive rotational failure 0.2342 610 170 10U 642859 6253490 Retrogressive rotational failure 0.236 545 3 10U 635265 6222327 Retrogressive rotational failure Pond location on slide Below headscarp (large, intact) Debris apron Next to small scarp (in debris) At base of headscarp (large, intact) Below large scarp (intact) Below small scarp (intact) Below small scarp (in debris) Just below small scarp near headscarp (large, intact) Below headscarp (large, intact) Below small scarp (intact) Below small scarp (intact) Just above small scarp (in debris) In debris apron below headscarp (large, intact) Next to small scarp (intact) and backtilt Below small scarp (intact) Base of headscarp (large, intact) Bottom of slide Below small scarp (intact) Just above large scarp (intact), near river Below small scarp (in debris) Base of headscarp (large, intact), next to small scarp (in debris) Above small scarp, below headscarp (large, intact) Below headscarp (large, intact) Above small scarp (in debris) Below small scarp (intact) Flat area midway down slide Between 2 small scarps (intact), by river Above small scarp (in debris) Base of headscarp (large, intact), above small scarp (intact) Below small scarp (intact) Body Above small scarp (intact) Below, to east of small scarp (in debris) 297 Dormant Abandoned Dormant Dormant Dormant Dormant Dormant Abandoned Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Abandoned Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Slide Activity Appendix 7 Landslide pond descriptions - Mapsheet 94A - cont'd Geomorphic location of pond Head Toe Body Body Body Body Body Head Body Body Body Body Body Body Toe Body Toe Body Toe Toe Body Head Body Toe Body Toe Toe Body Body Body Body Body Body Pond Pond Slide Size Slide UTMs Slide Type # # (ha) 0.2365 64 38 10U 652244 6236095 Retrogressive rotational failure 0.2384 538 3 10U 635265 6222327 Retrogressive rotational failure 0.2384 691 201 10U 655452 6222977 Retrogressive rotational failure 0.2407 608 168 10U 642871 6251602 Rotational failure 0.2466 5 59 10U 646337 6252998 Retrogressive rotational failure 0.249 63 38 10U 652244 6236095 Retrogressive rotational failure 0.2519 106 48 10U 641990 6244350 Rotational failure 0.2536 319 90 10U 584584 6239849 Retrogressive rotational failure 0.2545 109 48 10U 641990 6244350 Rotational failure 0.2559 619 175 10U 641263 6274322 Retrogressive rotational failure 0.2574 246 1 10U 631931 6221078 Retrogressive rotational failure 0.2578 475 126 10U 631665 6307193 Rotational failure 0.2581 236 1 10U 631931 6221078 Retrogressive rotational failure 0.2585 146 76 10U 633480 6244644 Retrogressive rotational failure 0.2613 770 227 10U 673362 6239998 Retrogressive rotational failure 0.2664 28 57 10U 645882 6252696 Retrogressive rotational failure 0.2759 155 62 10U 650179 6264258 Compound failure 0.2824 552 146 10U 635074 6225748 Multi-level rotational failure 0.2827 218 4 10U 637998 6220826 Multi-level rotational failure 0.2869 266 83 10U 611833 6238026 Retrogressive rotational failure 0.2874 43 71 10U 660889 6229983 Shallow retrogressive failure 0.2893 556 148 10U 635615 6217619 Retrogressive rotational failure 0.2906 304 87 10U 580867 6250762 Compound failure 0.2919 174 81 10U 673096 6235267 Retrogressive rotational failure 0.2934 203 14 10U 655144 6217752 Compound failure 0.3078 66 38 10U 652244 6236095 Retrogressive rotational failure 0.3181 452 122 10U 619929 6209531 Mobile flow 0.3193 248 1 10U 631931 6221078 Retrogressive rotational failure 0.3212 446 122 10U 619929 6209531 Mobile flow 0.3226 480 127 10U 632593 6304450 Rotational failure 0.3227 334 90 10U 584584 6239849 Retrogressive rotational failure 0.3239 427 114 10U 607383 6243249 Retrogressive rotational failure 0.3251 72 38 10U 652244 6236095 Retrogressive rotational failure Pond location on slide Toe Just below small scarp (intact) Bottom of slide Debris apron next to small scarp (intact) Below small scarp (in debris) Below small scarp (in debris) At base of headscarp (large, intact) Below headscarp (large, intact) Just above debris apron Below small scarp (in debris) Backtilt below small scarp (intact) Base of headscarp (small, intact) Below small scarp (intact) - near river Below small scarp (in debris) Base of headscarp (small, intact) Below headscarp (large, intact) Just below headscarp (small, intact) Above small scarp (intact) Below headscarp (large, intact), next to small scarp Above large scarp (intact) Just at base of headscarp (small, intact) Base of headscarp (small, intact) Within headscarp (small, intact) Below headscarp (large, intact) Within small scarp (in debris), just below headscarp (small, intact) Just below headscarp (large, intact) Centre of slide Above backtilt Near base of slide Below headscarp (small, intact) Just above small retrogressive scarp (intact) Base of headscarp (large, intact) Just below headscarp (large, intact) 298 Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Active Dormant Dormant Dormant Dormant Dormant Dormant Dormant Active Dormant Dormant Active Dormant Active Dormant Dormant Dormant Dormant Slide Activity Appendix 7 Landslide pond descriptions - Mapsheet 94A - cont'd Geomorphic location of pond Toe Body Toe Body Body Toe Body Head Body Toe Body Head Toe Body Head Body Head Body Body Body Head Head Head Body Head Head Body Body Body Body Body Head Head Pond Pond Slide Size Slide UTMs Slide Type # # (ha) 0.328 112 49 10U 641388 6245365 Multi-level rotational failure 0.3531 320 90 10U 584584 6239849 Retrogressive rotational failure 0.3538 234 1 10U 631931 6221078 Retrogressive rotational failure 0.3548 194 11 10U 656217 6219302 Retrogressive rotational failure 0.3553 695 10 10U 652580 6220750 Retrogressive rotational failure 0.3565 114 51 10U 642403 6248773 Rotational failure 0.3582 318 90 10U 584584 6239849 Retrogressive rotational failure 0.3589 239 1 10U 631931 6221078 Retrogressive rotational failure 0.3648 647 186 10U 655856 6238250 Retrogressive rotational failure 0.3731 633 182 10U 653926 6268718 Retrogressive rotational failure 0.3802 68 38 10U 652244 6236095 Retrogressive rotational failure 0.3815 245 1 10U 631931 6221078 Retrogressive rotational failure 0.3828 357 97 10U 597665 6245826 Rotational failure 0.3829 104 47 10U 640425 6241038 Retrogressive rotational failure 0.3898 532 1 10U 631931 6221078 Retrogressive rotational failure 0.397 739 211 10U 655836 6216390 Retrogressive rotational failure 0.4329 493 134 10U 634160 6291151 Rotational failure 0.436 694 159 10U 649510 6220866 Rotational failure 0.4523 270 83 10U 611833 6238026 Retrogressive rotational failure 0.4636 296 84 10U 612699 6237705 Retrogressive rotational failure 0.4648 130 66 10U 640009 6272311 Retrogressive rotational failure 0.4707 70 38 10U 652244 6236095 Retrogressive rotational failure 0.471 10 57 10U 645882 6252696 Retrogressive rotational failure 0.4942 9 59 10U 646337 6252998 Retrogressive rotational failure 0.506 652 189 10U 654871 6235759 Retrogressive rotational failure 0.51 134 70 10U 631913 6311160 Retrogressive rotational failure 0.515 232 6 10U 638745 6221151 Retrogressive rotational failure 0.519 178 18 10U 672960 6219166 Retrogressive rotational failure 0.525 186 10 10U 652580 6220750 Retrogressive rotational failure 0.527 600 48 10U 641990 6244350 Rotational failure 0.528 267 83 10U 611833 6238026 Retrogressive rotational failure 0.532 653 189 10U 654871 6235759 Retrogressive rotational failure 0.547 208 7 10U 637199 6218231 Retrogressive rotational failure Pond location on slide Below small scarp (in debris) Just below headscarp (large, intact) Above small scarp (intact) next to river Just above small scarp (in debris) Below headscarp (small, intact), above small scarp (in debris) In debris apron below headscarp (large, intact), next to south sidescarp Below headscarp (large, intact) Backtilt below headscarp (large, intact) Below small scarp (in debris), right by creek Above small scarp (intact) Above small scarp (intact) Backtilt below headscarp (large, intact) Just below headscarp (large, intact) Below headscarp (large, intact) Below small scarp (intact), by river Base of headscarp (large, intact) Bottom of slide Below small scarp (in debris) Below large scarp (intact) At base of small scarp (intact) Below headscarp, above small scarp (in debris) Just below headscarp (large, intact) Between 2 small scarps (in debris) Below small scarp (in debris) Below headscarp (large, intact) Below headscarp (small, intact) Just above small scarp (in debris) Above small scarp (in debris) Above small scarp (in debris), below headscarp (small, intact) Base of headscarp (large, intact) Above large scarp (intact) Below headscarp (large, intact) Below headscarp (east) - (small, intact) 299 Dormant Dormant Dormant Active Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Very active Dormant Active Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Slide Activity Appendix 7 Landslide pond descriptions - Mapsheet 94A - cont'd Geomorphic location of pond Body Head Toe Body Head Body Toe Head Toe Body Body Head Body Head Toe Head Toe Head Body Toe Body Head Body Body Body Head Body Body Head Body Body Body Body Pond Pond Slide Size Slide UTMs Slide Type # # (ha) 0.551 262 82 10U 564797 6216748 Rotational failure 0.552 71 38 10U 652244 6236095 Retrogressive rotational failure 0.565 214 9 10U 639115 6218246 Rotational failure 0.578 468 124 10U 621307 6287220 Rotational failure 0.582 185 10 10U 652580 6220750 Retrogressive rotational failure 0.582 69 38 10U 652244 6236095 Retrogressive rotational failure 0.6 297 85 10U 612971 6237140 Retrogressive rotational failure 0.601 148 76 10U 633480 6244644 Retrogressive rotational failure 0.606 494 135 10U 632986 6245453 Retrogressive rotational failure 0.608 353 95 10U 589923 6234772 Multi-level rotational failure 0.609 244 1 10U 631931 6221078 Retrogressive rotational failure 0.614 787 235 10U 685591 6226178 Multi-level rotational failure 0.617 253 1 10U 631931 6221078 Retrogressive rotational failure 0.617 8 59 10U 646337 6252998 Retrogressive rotational failure 0.627 237 1 10U 631931 6221078 Retrogressive rotational failure 0.637 627 179 10U 636025 6286564 Retrogressive rotational failure 0.64 477 127 10U 632593 6304450 Rotational failure 0.648 555 147 10U 636595 6219522 Multi-level rotational failure 0.661 235 1 10U 631931 6221078 Retrogressive rotational failure 0.668 271 83 10U 611833 6238026 Retrogressive rotational failure 0.672 295 84 10U 612699 6237705 Retrogressive rotational failure 0.675 333 90 10U 584584 6239849 Retrogressive rotational failure 0.758 184 10 10U 652580 6220750 Retrogressive rotational failure 0.815 183 10 10U 652580 6220750 Retrogressive rotational failure 0.858 0 15 10U 655647 6215120 Retrogressive rotational failure 0.874 623 177 10U 638157 6284497 Rotational failure 0.917 495 135 10U 632986 6245453 Retrogressive rotational failure 0.965 103 47 10U 640425 6241038 Retrogressive rotational failure 0.969 628 179 10U 636025 6286564 Retrogressive rotational failure 0.976 65 39 10U 651511 6236995 Rotational failure 0.989 536 3 10U 635265 6222327 Retrogressive rotational failure 1.025 78 38 10U 652244 6236095 Retrogressive rotational failure 1.062 561 150 10U 635070 6214869 Retrogressive rotational failure Pond location on slide At base of debris apron Below headscarp (large, intact) , above small scarp (in debris) Below small scarp (intact) Base of slide, near river Above small scarp (in debris), just below headscarp (backtilt - small, intact) Just below headscarp (large, intact) Below small scarp (intact) Below and within small scarp (in debris) Below small scarp (in debris) Below small scarp (intact) near river Backtilt below small scarp (intact) Base of large scarp (intact), in debris apron Above backtilt Just below headscarp (large, intact) Backtilt below headscarp (large, intact) Bottom of slide Below small scarp (intact) Debris apron Backtilt below small scarp (in debris) Below large scarp (intact) At base of small scarp (in debris) Just below headscarp (large, intact) Above small scarp (in debris), in backtilt below headscarp Above small scarp (intact) Just above small scarp (in debris), below headscarp (small, intact) Above/next to small scarp (intact) Below small scarp (in debris) Below small scarp (in debris) Bottom of slide Debris apron Below small scarp (intact) Toe Base of headscarp (small, intact) 300 Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Slide Activity Appendix 7 Landslide pond descriptions - Mapsheet 94A - cont'd Geomorphic location of pond Toe Head Toe Toe Head Head Toe Body Body Toe Body Toe Body Head Head Toe Body Toe Body Body Toe Head Head Body Head Body Body Toe Toe Toe Body Toe Head Pond Pond Slide Size Slide UTMs Slide Type # # (ha) 1.094 272 83 10U 611833 6238026 Retrogressive rotational failure 1.14 487 131 10U 632364 6300428 Rotational failure 1.22 105 48 10U 641990 6244350 Rotational failure 1.255 67 38 10U 652244 6236095 Retrogressive rotational failure 1.279 193 11 10U 656217 6219302 Retrogressive rotational failure 1.314 74 38 10U 652244 6236095 Retrogressive rotational failure 1.355 177 18 10U 672960 6219166 Retrogressive rotational failure 1.371 238 1 10U 631931 6221078 Retrogressive rotational failure 1.48 332 90 10U 584584 6239849 Retrogressive rotational failure 1.572 147 76 10U 633480 6244644 Retrogressive rotational failure 1.654 611 171 10U 641105 6253591 Rotational failure 2.376 73 44 10U 646758 6237400 Retrogressive rotational failure 3.227 629 179 10U 636025 6286564 Retrogressive rotational failure 5.89 207 7 10U 637199 6218231 Retrogressive rotational failure Pond location on slide At base of large scarp (intact) Middle of slide At base of headscarp (large, intact) Just below headscarp (large, intact) Below small scarp (in debris) Just below headscarp (large, intact) Below small scarp (in debris) Backtilt below headscarp (large, intact) Just below headscarp (large, intact) Just below headscarp (small, intact), just above small scarp (in debris) Bottom of slide, partly blocking creek Just below headscarp (large, intact), above small scarp (in debris) Bottom of slide Below headscarp (east) - (small, intact) 301 Dormant Dormant Dormant Dormant Active Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Dormant Slide Activity Appendix 7 Landslide pond descriptions - Mapsheet 94A - cont'd Geomorphic location of pond Body Body Body Head Body Head Toe Head Head Head Toe Body Toe Body Appendix 8 Pond size statistics by landslide type Slide type Compound failure Mobile flow Multi-level rotational Retrogressive rotational Rotational Shallow retrogressive Mean pond size (ha) 0.054 0.077 0.102 0.197 0.125 0.051 Minimum Maximum Median pond size pond size pond size (ha) (ha) (ha) 0.003 0.293 0.026 0.005 0.321 0.048 0.004 0.648 0.055 0.001 5.890 0.059 0.003 1.654 0.046 0.003 0.287 0.014 302 Standard Deviation of pond size (ha) 0.062 0.089 0.131 0.430 0.234 0.074 Appendix 9 Pond size by geomorphic location per slide type Ponds on head Landslide type Compound failure Mobile flow Multi-level rotational failure Retrogressive rotational failure Rotational failure Shallow retrogressive failure Ponds on body Landslide type Compound failure Mobile flow Multi-level rotational failure Retrogressive rotational failure Rotational failure Shallow retrogressive failure Ponds on toe Landslide type Compound failure Mobile flow Multi-level rotational failure Retrogressive rotational failure Rotational failure Shallow retrogressive failure Mean pond size (ha) Minimum pond size (ha) Maximum pond size (ha) Median pond size (ha) SD of pond size (ha) 0.072 0.053 0.078 0.281 0.082 0.064 0.003 0.025 0.012 0.007 0.004 0.003 0.293 0.109 0.181 1.572 0.436 0.287 0.022 0.033 0.081 0.131 0.065 0.009 0.100 0.030 0.047 0.366 0.090 0.112 0.050 0.085 0.104 0.192 0.123 0.025 0.007 0.005 0.007 0.001 0.003 0.003 0.293 0.321 0.060 5.890 1.654 0.068 0.033 0.046 0.058 0.062 0.042 0.007 0.046 0.100 0.099 0.489 0.222 0.025 0.050 0.000 0.109 0.150 0.159 0.072 0.004 0.000 0.004 0.002 0.007 0.059 0.201 0.000 0.648 0.358 1.654 0.081 0.017 0.000 0.044 0.041 0.047 0.075 0.058 0.000 0.160 0.355 0.310 0.009 303